Species in mammal family Ctenomyidae
Silky tuco-tuco (Ctenomys sericeus )
Ctenomyidae is a family of fossorial mammals in the order Rodentia and part of the Caviomorpha parvorder . Members of this family are called ctenomyids or tuco-tucos. They are found in southern South America, primarily in shrublands and grasslands , though some species can be found in forests, savannas , deserts, wetlands, and coastal areas. They range in size from the white-toothed tuco-tuco , at 11 cm (4 in) plus a 7 cm (3 in) tail, to Conover's tuco-tuco , at 33 cm (13 in) plus a 11 cm (4 in) tail. Ctenomyids are herbivores and eat a wide variety of vegetation.[ 1] No ctenomyids have population estimates, but ten species are categorized as an endangered species , and three—Reig's tuco-tuco , Roig's tuco-tuco , and the social tuco-tuco —are categorized as critically endangered .
The 59 extant species of Ctenomyidae are all contained in a single genus , Ctenomys . A few extinct prehistoric ctenomyid species have been discovered, though due to ongoing research and discoveries, the exact number and categorization is not fixed.[ 2]
Conventions
The author citation for the species or genus is given after the scientific name; parentheses around the author citation indicate that this was not the original taxonomic placement. Conservation status codes listed follow the International Union for Conservation of Nature (IUCN) Red List of Threatened Species . Range maps are provided wherever possible; if a range map is not available, a description of the ctenomyid's range is provided. Ranges are based on the IUCN Red List for that species unless otherwise noted.
Classification
Ctenomyidae is a family consisting of 59 species in a single genus , Ctenomys .
Ctenomyids
The following classification is based on the taxonomy described by the reference work Mammal Species of the World (2005), with augmentation by generally accepted proposals made since using molecular phylogenetic analysis , as supported by both the IUCN and the American Society of Mammalogists .[ 3]
Genus Ctenomys – Blainville , 1826 – 59 species
Common name
Scientific name and subspecies
Range
Size and ecology
IUCN status and estimated population
Argentine tuco-tuco
C. argentinus Berry & Contreras , 1982
Northern Argentina
Size : 17–18 cm (7 in) long, plus 7–9 cm (3–4 in) tail[ 4] Habitat : Shrubland and grassland[ 5]
NT
Unknown [ 5]
Azara's tuco-tuco
C. azarae Thomas , 1903
Central Argentina
Size : 16–20 cm (6–8 in) long, plus 6–9 cm (2–4 in) tail[ 6] Habitat : Shrubland[ 7]
EN
Unknown [ 7]
Berg's tuco-tuco
C. bergi Thomas , 1902
Central Argentina
Size : 13–17 cm (5–7 in) long, plus about 7 cm (3 in) tail[ 8] Habitat : Savanna and grassland[ 9]
EN
Unknown [ 9]
Bolivian tuco-tuco
C. boliviensis Waterhouse , 1848
C. b. boliviensis
C. b. nattereri
Bolivia and western Brazil
Size : 18–28 cm (7–11 in) long, plus 7–10 cm (3–4 in) tail[ 10] Habitat : Savanna[ 11]
LC
Unknown [ 11]
Bonetto's tuco-tuco
C. bonettoi Berry & Contreras , 1982
Northern Argentina
Size : 17–19 cm (7 in) long, plus 6–8 cm (2–3 in) tail[ 8] Habitat : Shrubland[ 12]
EN
Unknown [ 12]
Brazilian tuco-tuco
C. brasiliensis Blainville , 1826
Uruguay
Size : Unknown[ 6] Habitat : Unknown[ 13]
DD
Unknown [ 13]
Catamarca tuco-tuco
C. knighti Thomas , 1919
Northern Argentina
Size : About 20 cm (8 in) long, plus about 8 cm (3 in) tail[ 8] Habitat : Grassland[ 14]
DD
Unknown [ 14]
Chacoan tuco-tuco
C. dorsalis Thomas , 1900
Paraguay and Bolivia
Size : About 16 cm (6 in) long, plus about 5 cm (2 in) tail[ 15] Habitat : Savanna[ 16]
DD
Unknown [ 16]
Colburn's tuco-tuco
C. colburni Allen , 1903
Southern Argentina
Size : Unknown[ 4] Habitat : Grassland[ 17]
DD
Unknown [ 17]
Collared tuco-tuco
C. torquatus Lichtenstein , 1830
Uruguay and southern Brazil
Size : 15–23 cm (6–9 in) long, plus 5–9 cm (2–4 in) tail[ 15] Habitat : Forest and shrubland[ 18]
LC
Unknown [ 18]
Conover's tuco-tuco
C. conoveri Osgood , 1946
Paraguay and Bolivia
Size : 24–33 cm (9–13 in) long, plus 9–14 cm (4–6 in) tail[ 19] Habitat : Shrubland and grassland[ 20]
LC
Unknown [ 20]
Coyhaique tuco-tuco
C. coyhaiquensis Kelt & Gallardo , 1994
Southern Argentina and southern Chile
Size : Unknown length, plus 5–9 cm (2–4 in) tail[ 4] Habitat : Shrubland[ 21]
DD
Unknown [ 21]
D'Orbigny's tuco-tuco
C. dorbignyi Contreras & Contreras , 1984
Northern Argentina
Size : 19–22 cm (7–9 in) long, plus 8–10 cm (3–4 in) tail[ 6] Habitat : Shrubland and grassland[ 22]
NT
Unknown [ 22]
Emily's tuco-tuco
C. emilianus St. Leger , 1926
Western Argentina
Size : About 21 cm (8 in) long, plus about 9 cm (4 in) tail[ 10] Habitat : Desert[ 23]
LC
Unknown [ 23]
Famatina tuco-tuco
C. famosus Thomas , 1920
Northwestern Argentina
Size : About 16 cm (6 in) long, plus about 7 cm (3 in) tail[ 15] Habitat : Unknown[ 24]
DD
Unknown [ 24]
Flamarion's tuco-tuco
C. flamarioni Travi , 1981
Uruguay and southern Brazil
Size : About 25 cm (10 in) long, plus about 7 cm (3 in) tail[ 6] Habitat : Coastal marine[ 25]
EN
Unknown [ 25]
Foch's tuco-tuco
C. fochi Thomas , 1919
Northern Argentina
Size : About 16 cm (6 in) long, plus about 8 cm (3 in) tail[ 8] Habitat : Unknown[ 26]
DD
Unknown [ 26]
Furtive tuco-tuco
C. occultus Thomas , 1920
Northern Argentina
Size : 13–16 cm (5–6 in) long, plus tail[ 4] Habitat : Unknown[ 27]
EN
Unknown [ 27]
Goodfellow's tuco-tuco
C. goodfellowi Thomas , 1921
Bolivia
Size : 18–28 cm (7–11 in) long, plus 7–10 cm (3–4 in) tail[ 10] Habitat : Forest[ 28]
LC
Unknown [ 28]
Goya tuco-tuco
C. perrensi Thomas , 1898
Northeastern Argentina
Size : About 20 cm (8 in) long, plus about 7 cm (3 in) tail[ 6] Habitat : Unknown[ 29]
LC
Unknown [ 29]
Haig's tuco-tuco
C. haigi Thomas , 1917
C. h. haigi
C. h. lentulus
Western Argentina
Size : 17–20 cm (7–8 in) long, plus 8–9 cm (3–4 in) tail[ 4] Habitat : Forest, grassland, and inland wetlands[ 30]
LC
Unknown [ 30]
Highland tuco-tuco
C. opimus Wagner , 1848
C. o. luteolus
C. o. nigriceps
C. o. opimus
Bolivia and northwestern Argentina
Size : 19–25 cm (7–10 in) long, plus 7–10 cm (3–4 in) tail[ 15] Habitat : Grassland[ 31]
LC
Unknown [ 31]
Ibicui tuco-tuco
C. ibicui de Freitas , Fernandes , Fornel , Roratto , 2012
Southern Brazil
Size : About 16 cm (6 in) long, plus about 8 cm (3 in) tail[ 15] Habitat : Grassland[ 32]
NT
Unknown [ 32]
Jujuy tuco-tuco
C. juris Thomas , 1920
Northern Argentina
Size : About 18 cm (7 in) long, plus about 7 cm (3 in) tail[ 8] Habitat : Unknown[ 33]
DD
Unknown [ 33]
Lago Blanco tuco-tuco
C. fodax Thomas , 1910
Southern Argentina and southern Chile
Size : About 26 cm (10 in) long, plus about 10 cm (4 in) tail[ 4] Habitat : Grassland[ 34]
DD
Unknown [ 34]
Lami tuco-tuco
C. lami Freitas , 2001
Southern Brazil
Size : 16–22 cm (6–9 in) long, plus 6–10 cm (2–4 in) tail[ 15] Habitat : Coastal marine[ 35]
VU
Unknown [ 35]
Lewis's tuco-tuco
C. lewisi Thomas , 1926
Southern Bolivia
Size : 20–22 cm (8–9 in) long, plus 6–8 cm (2–3 in) tail[ 4] Habitat : Grassland[ 36]
LC
Unknown [ 36]
Magellanic tuco-tuco
C. magellanicus Bennett , 1836
C. m. dicki
C. m. fueginus
C. m. magellanicus
C. m. osgoodi
Southern Argentina and southern Chile
Size : Unknown[ 4] Habitat : Grassland[ 37]
LC
Unknown [ 37]
Maule tuco-tuco
C. maulinus Philippi , 1872
C. m. brunneus
C. m. maulinus
Central Chile
Size : Unknown[ 6] Habitat : Grassland[ 38]
LC
Unknown [ 38]
Mendoza tuco-tuco
C. mendocinus Philippi , 1869
Central Argentina
Size : 16–20 cm (6–8 in) long, plus 6–9 cm (2–4 in) tail[ 6] Habitat : Shrubland, grassland, and inland wetlands[ 39]
LC
Unknown [ 39]
Mottled tuco-tuco
C. latro Thomas , 1918
Northern Argentina
Size : 16–18 cm (6–7 in) long, plus tail[ 4] Habitat : Shrubland and grassland[ 40]
EN
Unknown [ 40]
Pearson's tuco-tuco
C. pearsoni Lessa & Langguth , 1983
Southern Uruguay
Size : 17–20 cm (7–8 in) long, plus 7–9 cm (3–4 in) tail[ 6] Habitat : Shrubland, grassland, and coastal marine[ 41]
NT
Unknown [ 41]
Peruvian tuco-tuco
C. peruanus Sanborn & Pearson , 1947
Southern Peru
Size : About 22 cm (9 in) long, plus about 9 cm (4 in) tail[ 8] Habitat : Grassland[ 42]
LC
Unknown [ 42]
Pilar tuco-tuco
C. pilarensis Contreras , 1993
Northern Argentina and Paraguay
Size : 16–21 cm (6–8 in) long, plus tail[ 4] Habitat : Grassland[ 43]
EN
Unknown [ 43]
Porteous's tuco-tuco
C. porteousi Thomas , 1916
Central Argentina
Size : 16–20 cm (6–8 in) long, plus 6–9 cm (2–4 in) tail[ 6] Habitat : Grassland[ 44]
NT
Unknown [ 44]
Pundt's tuco-tuco
C. pundti Nehring , 1900
Central Argentina
Size : About 17 cm (7 in) long, plus about 4 cm (2 in) tail[ 6] Habitat : Grassland[ 45]
EN
Unknown [ 45]
Puntilla tuco-tuco
C. coludo Thomas , 1920
Northwestern Argentina
Size : About 20 cm (8 in) long, plus about 10 cm (4 in) tail[ 15] Habitat : Unknown[ 46]
DD
Unknown [ 46]
Reddish tuco-tuco
C. frater Thomas , 1902
C. f. frater
C. f. mordosus
Bolivia and northern Argentina
Size : 17–20 cm (7–8 in) long, plus 5–9 cm (2–4 in) tail[ 4] Habitat : Grassland[ 47]
LC
Unknown [ 47]
Reig's tuco-tuco
C. osvaldoreigi Contreras , 1995
Central Argentina
Size : 23–25 cm (9–10 in) long, plus tail[ 8] Habitat : Grassland[ 48]
CR
Unknown [ 48]
Rio Negro tuco-tuco
C. rionegrensis Langguth & Abella , 1970
Eastern Argentina and Uruguay
Size : 16–19 cm (6–7 in) long, plus 6–9 cm (2–4 in) tail[ 6] Habitat : Desert and coastal marine[ 49]
EN
Unknown [ 49]
Robust tuco-tuco
C. tuconax Thomas , 1925
Northern Argentina
Size : Unknown[ 10] Habitat : Savanna, shrubland, and grassland[ 50]
DD
Unknown [ 50]
Roig's tuco-tuco
C. roigi Contreras , 1988
Northern Argentina
Size : 17–20 cm (7–8 in) long, plus 7–10 cm (3–4 in) tail[ 6] Habitat : Grassland, inland wetlands, and other[ 51]
CR
Unknown [ 51]
Salta tuco-tuco
C. saltarius Thomas , 1912
Northern Argentina
Size : About 20 cm (8 in) long, plus about 9 cm (4 in) tail[ 15] Habitat : Desert[ 52]
DD
Unknown [ 52]
San Juan tuco-tuco
C. johannis Thomas , 1921
Western Argentina
Size : About 20 cm (8 in) long, plus about 10 cm (4 in) tail[ 15] Habitat : Unknown[ 53]
DD
Unknown [ 53]
San Luis tuco-tuco
C. pontifex Thomas , 1918
Central Argentina and central Chile
Size : About 18 cm (7 in) long, plus about 8 cm (3 in) tail[ 8] Habitat : Unknown[ 54]
DD
Unknown [ 54]
Scaglia's tuco-tuco
C. scagliai Contreras , 1999
Northern Argentina
Size : About 20 cm (8 in) long, plus about 9 cm (4 in) tail[ 15] Habitat : Unknown[ 55]
DD
Unknown [ 55]
Sierra Tontal tuco-tuco
C. tulduco Thomas , 1921
Western Argentina
Size : About 19 cm (7 in) long, plus about 7 cm (3 in) tail[ 15] Habitat : Unknown[ 56]
DD
Unknown [ 56]
Silky tuco-tuco
C. sericeus Allen , 1903
Southern Argentina and southern Chile
Size : Unknown length, plus 5–9 cm (2–4 in) tail[ 4] Habitat : Forest[ 57]
DD
Unknown [ 57]
Social tuco-tuco
C. sociabilis Pearson & Christie , 1985
Central Argentina and central Chile
Size : 16–25 cm (6–10 in) long, plus 6–8 cm (2–3 in) tail[ 10] Habitat : Savanna and inland wetlands[ 58]
CR
Unknown [ 58]
Southern tuco-tuco
C. australis Rusconi , 1934
Eastern Argentina
Size : About 21 cm (8 in) long, plus 9–11 cm (4 in) tail[ 6] Habitat : Coastal marine[ 59]
EN
Unknown [ 59]
Steinbach's tuco-tuco
C. steinbachi Thomas , 1907
Bolivia
Size : 23–25 cm (9–10 in) long, plus tail[ 10] Habitat : Savanna[ 60]
LC
Unknown [ 60]
Strong tuco-tuco
C. validus Contreras , Roig , & Suzarte , 1977
Central Argentina
Size : Unknown[ 15] Habitat : Unknown[ 61]
DD
Unknown [ 61]
Talas tuco-tuco
C. talarum Thomas , 1898
C. t. antonii
C. t. occidentalis
C. t. recessus
C. t. talarum
Central and eastern Argentina
Size : 15–18 cm (6–7 in) long, plus 5–8 cm (2–3 in) tail[ 6] Habitat : Grassland and coastal marine[ 62]
LC
Unknown [ 62]
Tawny tuco-tuco
C. fulvus Philippi , 1860
C. f. fulvus
C. f. robustus
Western South America
Size : Unknown[ 15] Habitat : Forest and desert[ 63]
DD
Unknown [ 63]
Tiny tuco-tuco
C. minutus Nehring , 1887
C. m. bicolor
C. m. minutus
Southern Brazil
Size : About 7 cm (3 in) long, plus tail[ 15] Habitat : Shrubland and grassland[ 64]
DD
Unknown [ 64]
Tucuman tuco-tuco
C. tucumanus Thomas , 1900
Northern Argentina
Size : About 17 cm (7 in) long, plus about 7 cm (3 in) tail[ 4] Habitat : Forest[ 65]
DD
Unknown [ 65]
Vipos tuco-tuco
C. viperinus Thomas , 1926
Northern Argentina
Size : About 21 cm (8 in) long, plus about 8 cm (3 in) tail[ 8] Habitat : Unknown[ 66]
DD
Unknown [ 66]
White-toothed tuco-tuco
C. leucodon Waterhouse , 1848
Western Bolivia and southern Peru
Size : 11–18 cm (4–7 in) long, plus 7–9 cm (3–4 in) tail[ 4] Habitat : Shrubland and grassland[ 67]
LC
Unknown [ 67]
Yolanda's tuco-tuco
C. yolandae Contreras & Berry , 1984
Northern Argentina
Size : Unknown[ 8] Habitat : Unknown[ 68]
DD
Unknown [ 68]
References
^ Nowak , p. 1680
^ "Fossilworks: Ctenomyidae" . Paleobiology Database . University of Wisconsin–Madison . Archived from the original on December 4, 2024. Retrieved June 30, 2025 .
^ Wilson, Reeder , pp. 1560-1569
^ a b c d e f g h i j k l m n Chernasky; Motis; Burgin , p. 363
^ a b Bidau, C. J. (2019). "Ctenomys argentinus " . IUCN Red List of Threatened Species . 2019 : e.T5795A22191874. doi :10.2305/IUCN.UK.2019-3.RLTS.T5795A22191874.en .
^ a b c d e f g h i j k l m n Chernasky; Motis; Burgin , p. 362
^ a b Bidau, C. J. (2019). "Ctenomys azarae " . IUCN Red List of Threatened Species . 2019 : e.T5797A22192079. doi :10.2305/IUCN.UK.2019-3.RLTS.T5797A22192079.en .
^ a b c d e f g h i j Chernasky; Motis; Burgin , p. 364
^ a b Bidau, C. J. (2019). "Ctenomys bergi " . IUCN Red List of Threatened Species . 2019 : e.T136687A22196510. doi :10.2305/IUCN.UK.2019-3.RLTS.T136687A22196510.en .
^ a b c d e f Chernasky; Motis; Burgin , p. 360
^ a b Dunnum, J.; Bernal, N. (2017) [errata version of 2016 assessment]. "Ctenomys boliviensis " . IUCN Red List of Threatened Species . 2016 : e.T5798A115079917. doi :10.2305/IUCN.UK.2016-3.RLTS.T5798A22192171.en .
^ a b Bidau, C. J. (2018). "Ctenomys bonettoi " . IUCN Red List of Threatened Species . 2018 : e.T5799A22192271. doi :10.2305/IUCN.UK.2018-1.RLTS.T5799A22192271.en .
^ a b Ojeda, R. (2019). "Ctenomys brasiliensis " . IUCN Red List of Threatened Species . 2019 : e.T5800A22192465. doi :10.2305/IUCN.UK.2019-1.RLTS.T5800A22192465.en .
^ a b Ojeda, R. (2019). "Ctenomys knighti " . IUCN Red List of Threatened Species . 2019 : e.T5808A22193333. doi :10.2305/IUCN.UK.2019-1.RLTS.T5808A22193333.en .
^ a b c d e f g h i j k l m n Chernasky; Motis; Burgin , p. 361
^ a b Ojeda, R. (2019). "Ctenomys dorsalis " . IUCN Red List of Threatened Species . 2019 : e.T5803A22192708. doi :10.2305/IUCN.UK.2019-1.RLTS.T5803A22192708.en .
^ a b Bidau, C. J. (2019). "Ctenomys colburni " . IUCN Red List of Threatened Species . 2019 : e.T5801A22192538. doi :10.2305/IUCN.UK.2019-1.RLTS.T5801A22192538.en .
^ a b Bidau, C. (2017) [errata version of 2016 assessment]. "Ctenomys torquatus " . IUCN Red List of Threatened Species . 2016 : e.T5829A115080157. doi :10.2305/IUCN.UK.2016-3.RLTS.T5829A78319489.en .
^ Brashear, Katie (2004). "Ctenomys conoveri " . Animal Diversity Web . University of Michigan . Retrieved June 30, 2025 .
^ a b Dunnum, J.; Bernal, N. (2016). "Ctenomys conoveri " . IUCN Red List of Threatened Species . 2016 : e.T5802A22192609. doi :10.2305/IUCN.UK.2016-2.RLTS.T5802A22192609.en .
^ a b Ojeda, R. (2019). "Ctenomys coyhaiquensis " . IUCN Red List of Threatened Species . 2019 : e.T136295A22196080. doi :10.2305/IUCN.UK.2019-1.RLTS.T136295A22196080.en .
^ a b Bidau, C. J. (2019). "Ctenomys dorbignyi " . IUCN Red List of Threatened Species . 2019 : e.T136234A22192383. doi :10.2305/IUCN.UK.2019-3.RLTS.T136234A22192383.en .
^ a b Bidau, C. J.; Ojeda, R. (2019). "Ctenomys emilianus " . IUCN Red List of Threatened Species . 2019 : e.T5804A22192803. doi :10.2305/IUCN.UK.2019-3.RLTS.T5804A22192803.en .
^ a b Bidau, C. J. (2019). "Ctenomys famosus " . IUCN Red List of Threatened Species . 2019 : e.T136588A22197042. doi :10.2305/IUCN.UK.2019-1.RLTS.T136588A22197042.en .
^ a b de Frietas, T.; Weksler, M.; Catzeflis, F.; Percequillo, A. (2019). "Ctenomys flamarioni " . IUCN Red List of Threatened Species . 2019 : e.T136464A22196926. doi :10.2305/IUCN.UK.2019-3.RLTS.T136464A22196926.en .
^ a b Ojeda, R. (2019). "Ctenomys fochi " . IUCN Red List of Threatened Species . 2019 : e.T136499A22193166. doi :10.2305/IUCN.UK.2019-1.RLTS.T136499A22193166.en .
^ a b Bidau, C. J. (2018). "Ctenomys occultus " . IUCN Red List of Threatened Species . 2018 : e.T5817A22194328. doi :10.2305/IUCN.UK.2018-1.RLTS.T5817A22194328.en .
^ a b Roach, N.; Naylor, L. (2016). "Ctenomys goodfellowi " . IUCN Red List of Threatened Species . 2016 : e.T136607A22196835. doi :10.2305/IUCN.UK.2016-2.RLTS.T136607A22196835.en .
^ a b Bidau, C. (2016). "Ctenomys perrensi " . IUCN Red List of Threatened Species . 2016 : e.T5820A22194434. doi :10.2305/IUCN.UK.2016-2.RLTS.T5820A22194434.en .
^ a b Bidau, C. (2016). "Ctenomys haigi " . IUCN Red List of Threatened Species . 2016 : e.T5807A22193246. doi :10.2305/IUCN.UK.2016-2.RLTS.T5807A22193246.en .
^ a b Dunnum, J.; Vargas, J.; Bernal, N.; Zeballos, N.; Vivar, E. (2016). "Ctenomys opimus " . IUCN Red List of Threatened Species . 2016 : e.T5818A22194246. doi :10.2305/IUCN.UK.2016-2.RLTS.T5818A22194246.en .
^ a b Hurtado Materon, M. A.; de Frietas, T.; Lawing, M.; Gonçalves, G. L.; Fornel, R. (2024). "Ctenomys ibicuiensis " . IUCN Red List of Threatened Species . 2024 : e.T45958852A259145694. doi :10.2305/IUCN.UK.2024-2.RLTS.T45958852A259145694.en .
^ a b Ojeda, R. (2019). "Ctenomys juris " . IUCN Red List of Threatened Species . 2019 : e.T136556A22196428. doi :10.2305/IUCN.UK.2019-1.RLTS.T136556A22196428.en .
^ a b Ojeda, R. (2019). "Ctenomys fodax " . IUCN Red List of Threatened Species . 2019 : e.T136601A22196762. doi :10.2305/IUCN.UK.2019-1.RLTS.T136601A22196762.en .
^ a b Bidau, C. J. (2018). "Ctenomys lami " . IUCN Red List of Threatened Species . 2018 : e.T136567A22196245. doi :10.2305/IUCN.UK.2018-1.RLTS.T136567A22196245.en .
^ a b Roach, N.; Naylor, L. (2016). "Ctenomys lewisi " . IUCN Red List of Threatened Species . 2016 : e.T5811A22193810. doi :10.2305/IUCN.UK.2016-2.RLTS.T5811A22193810.en .
^ a b Bidau, C. J. (2019). "Ctenomys magellanicus " . IUCN Red List of Threatened Species . 2019 : e.T5812A22193726. doi :10.2305/IUCN.UK.2019-1.RLTS.T5812A22193726.en .
^ a b Bidau, C. (2016). "Ctenomys maulinus " . IUCN Red List of Threatened Species . 2016 : e.T5813A22193961. doi :10.2305/IUCN.UK.2016-2.RLTS.T5813A22193961.en .
^ a b Bidau, Ojeda; R. (2017) [errata version of 2016 assessment]. "Ctenomys mendocinus " . IUCN Red List of Threatened Species . 2016 : e.T5814A115080043. doi :10.2305/IUCN.UK.2016-3.RLTS.T5814A22193878.en .
^ a b Bidau, C. J. (2019). "Ctenomys latro " . IUCN Red List of Threatened Species . 2019 : e.T5809A22193629. doi :10.2305/IUCN.UK.2019-3.RLTS.T5809A22193629.en .
^ a b Bidau, C. J. (2019). "Ctenomys pearsoni " . IUCN Red List of Threatened Species . 2019 : e.T5819A22194513. doi :10.2305/IUCN.UK.2019-3.RLTS.T5819A22194513.en .
^ a b Zeballos, H.; Vivar, E. (2016). "Ctenomys peruanus " . IUCN Red List of Threatened Species . 2016 : e.T5821A22194718. doi :10.2305/IUCN.UK.2016-2.RLTS.T5821A22194718.en .
^ a b Bidau, C. J. (2019). "Ctenomys pilarensis " . IUCN Red List of Threatened Species . 2019 : e.T136289A22195989. doi :10.2305/IUCN.UK.2019-3.RLTS.T136289A22195989.en .
^ a b Bidau, C. J. (2020). "Ctenomys porteousi " . IUCN Red List of Threatened Species . 2020 : e.T5823A22194905. doi :10.2305/IUCN.UK.2020-3.RLTS.T5823A22194905.en .
^ a b Bidau, C. J. (2019). "Ctenomys pundti " . IUCN Red List of Threatened Species . 2019 : e.T136370A22194037. doi :10.2305/IUCN.UK.2019-3.RLTS.T136370A22194037.en .
^ a b Bidau, C. J.; Ojeda, R. (2019). "Ctenomys coludo " . IUCN Red List of Threatened Species . 2019 : e.T136308A22196340. doi :10.2305/IUCN.UK.2019-1.RLTS.T136308A22196340.en .
^ a b Vivar, E. (2018) [errata version of 2017 assessment]. "Ctenomys frater " . IUCN Red List of Threatened Species . 2017 : e.T115553730A123796865. doi :10.2305/IUCN.UK.2017-3.RLTS.T115553730A115553734.en .
^ a b Bidau, C. J. (2018). "Ctenomys osvaldoreigi " . IUCN Red List of Threatened Species . 2018 : e.T136390A22195057. doi :10.2305/IUCN.UK.2018-1.RLTS.T136390A22195057.en .
^ a b Bidau, C. J. (2018). "Ctenomys rionegrensis " . IUCN Red List of Threatened Species . 2018 : e.T136635A22193418. doi :10.2305/IUCN.UK.2018-1.RLTS.T136635A22193418.en .
^ a b Ojeda, R. (2019). "Ctenomys tuconax " . IUCN Red List of Threatened Species . 2019 : e.T5830A22195740. doi :10.2305/IUCN.UK.2019-1.RLTS.T5830A22195740.en .
^ a b Bidau, C. J. (2018). "Ctenomys roigi " . IUCN Red List of Threatened Species . 2018 : e.T136633A22193077. doi :10.2305/IUCN.UK.2018-1.RLTS.T136633A22193077.en .
^ a b Ojeda, R. (2019). "Ctenomys saltarius " . IUCN Red List of Threatened Species . 2019 : e.T5824A22194818. doi :10.2305/IUCN.UK.2019-1.RLTS.T5824A22194818.en .
^ a b Bidau, C. J. (2019). "Ctenomys johannis " . IUCN Red List of Threatened Species . 2019 : e.T136281A22195842. doi :10.2305/IUCN.UK.2019-1.RLTS.T136281A22195842.en .
^ a b Ojeda, R. (2019). "Ctenomys pontifex " . IUCN Red List of Threatened Species . 2019 : e.T5822A22194645. doi :10.2305/IUCN.UK.2019-1.RLTS.T5822A22194645.en .
^ a b Ojeda, R. (2019). "Ctenomys scagliai " . IUCN Red List of Threatened Species . 2019 : e.T136568A22196158. doi :10.2305/IUCN.UK.2019-1.RLTS.T136568A22196158.en .
^ a b Bidau, C. J. (2019). "Ctenomys tulduco " . IUCN Red List of Threatened Species . 2019 : e.T136826A22197188. doi :10.2305/IUCN.UK.2019-1.RLTS.T136826A22197188.en .
^ a b Bidau, C. J. (2019). "Ctenomys sericeus " . IUCN Red List of Threatened Species . 2019 : e.T5825A22195256. doi :10.2305/IUCN.UK.2019-1.RLTS.T5825A22195256.en .
^ a b Bidau, C. J. (2018). "Ctenomys sociabilis " . IUCN Red List of Threatened Species . 2018 : e.T5826A22195323. doi :10.2305/IUCN.UK.2018-1.RLTS.T5826A22195323.en .
^ a b Bidau, C. J. (2018). "Ctenomys australis " . IUCN Red List of Threatened Species . 2018 : e.T5796A78319377. doi :10.2305/IUCN.UK.2018-1.RLTS.T5796A78319377.en .
^ a b Roach, N.; Naylor, L. (2016). "Ctenomys steinbachi " . IUCN Red List of Threatened Species . 2016 : e.T5827A22194994. doi :10.2305/IUCN.UK.2016-2.RLTS.T5827A22194994.en .
^ a b Ojeda, R. (2019). "Ctenomys validus " . IUCN Red List of Threatened Species . 2019 : e.T5832A22195532. doi :10.2305/IUCN.UK.2019-1.RLTS.T5832A22195532.en .
^ a b Bidau, C. (2016). "Ctenomys talarum " . IUCN Red List of Threatened Species . 2016 : e.T5828A22195175. doi :10.2305/IUCN.UK.2016-2.RLTS.T5828A22195175.en .
^ a b Bidau, C. J.; Ojeda, R. (2019). "Ctenomys fulvus " . IUCN Red List of Threatened Species . 2019 : e.T5806A22192983. doi :10.2305/IUCN.UK.2019-1.RLTS.T5806A22192983.en .
^ a b Dunnum, J.; Bernal, N. (2019). "Ctenomys minutus " . IUCN Red List of Threatened Species . 2019 : e.T5815A22194130. doi :10.2305/IUCN.UK.2019-1.RLTS.T5815A22194130.en .
^ a b Ojeda, R. (2019). "Ctenomys tucumanus " . IUCN Red List of Threatened Species . 2019 : e.T5831A22195435. doi :10.2305/IUCN.UK.2019-1.RLTS.T5831A22195435.en .
^ a b Bidau, C. J. (2018). "Ctenomys viperinus " . IUCN Red List of Threatened Species . 2018 : e.T136758A22195904. doi :10.2305/IUCN.UK.2018-2.RLTS.T136758A22195904.en .
^ a b Zeballos, H.; Vivar, E. (2016). "Ctenomys leucodon " . IUCN Red List of Threatened Species . 2016 : e.T5810A22193526. doi :10.2305/IUCN.UK.2016-2.RLTS.T5810A22193526.en .
^ a b Bidau, C. J.; Ojeda, R. (2019). "Ctenomys yolandae " . IUCN Red List of Threatened Species . 2019 : e.T136681A22196679. doi :10.2305/IUCN.UK.2019-1.RLTS.T136681A22196679.en .
Sources