Плавци (породица инсеката)
Плавци (Lycaenidae)
Phengaris arion - пегави мравник
Дугорепи селац (Lampides boeticus)
Научна класификација
Царство:
Тип:
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(нерангирано):
Натпородица:
Породица:
Lycaenidae
Потпородице
Плавци [ 1] (лат. Lycaenidae) су велика породица дневних лептира , претежно плаве, наранџасте или смеђе боје горње стране крила. Заступљени су великим бројем врста. У Европи живи 170 врста плаваца, а у Србији 55 врста. Породица је подељена на више потпородица.
Lycaenidae је друга по величини породица лептира (иза Nymphalidae , четкастих лептира), са преко 6.000 врста широм света,[ 2] чији се чланови називају и лептири са паучинастим крилима . Они чине око 30% познатих врста лептира.
Опис, храна и животни циклус
Одрасле јединке су мале, обично испод 5 cm, и јарко обојене, понекад са металним сјајем. Код ове породице лептира најизраженији је полни диморфизам . Најчешће су женке смеђе и неупадљиве боје.
Ларве су често спљоштене, а не цилиндричне, са жлездама које могу производити секрет који привлаче и потчињавају мраве . Њихове кутикуле имају тенденцију да буду задебљане. Неке ларве су способне да производе вибрације и ниске звукове који се преносе кроз супстрате у којима живе. Оне користе ове звукове за комуникацију са мравима.[ 3] [ 4]
Одрасли појединци често имају длакаве репове налик на антене заједно са црно-белим прстенастим (прстенастим) изгледом. Многе врсте такође имају тачку у дну репа, а неке се окрећу по слетању како би збуниле потенцијалне предаторе да препознају праву оријентацију главе. Ово узрокује да предатори приђу са праве главе, што резултира раним визуелним откривањем.[ 5]
Мрав који чува ларву плавца - Lycaenidae
Lycaenidae су разнолики у погледу својих прехрамбених навика и осим фитофагије , неки од њих су ентомофагни , хране се лисним вашима , љускавим инсектима и ларвама мрава. Неки ликениди чак искоришћавају своју повезаност са мравима наводећи мраве да их хране регургитацијом, процесом који се назива трофалакса . Нису свим ликенидним лептирима потребни мрави, али око 75% врста се повезује са мравима,[ 3] однос који се назива мирмекофилија . Ове асоцијације могу бити узајамне, паразитске или предаторске у зависности од врсте.
Код неких врста, ларве посећују и штите мрави док се хране биљком домаћином, а мрави од њих добијају медну росу богату шећером током целог живота ларве, а код неких врста током стадијума кукуљице. Код других врста, само првих неколико фаза се проводи на биљци, а остатак животног века ларве проводи се као грабежљивац у гнезду мрава. Ларва постаје паразит, који се храни регургитацијом мрава, или предатор на ларвама мрава.[ 3] Гусенице се развијау унутар гнезда мрава и мрави настављају да брину о луткама. Непосредно пре изласка одраслих јединки, крила лептира унутар кућишта кукуљице се одвајају од ње, а лутка постаје сребрнаста. Одрасли лептир излази из кукуљице након три до четири недеље, још увек унутар гнезда мрава. Лептир мора да испузи из мрављег гнезда пре него што може да прошири крила.
Неколико еволуционих адаптација омогућава ове асоцијације, укључујући мале жлезде на кожи гусеница које се називају „органи поре куполе“. Гусенице многих врста имају жлезду на седмом трбушном сегменту која производи медљику и назива се „дорзална нектарна жлезда“ (она је позната и као „дошљачка жлезда“). На осмом трбушном сегменту налази се орган који се може окретати, назван „пипкасти орган”, цилиндричан је са прстеном шиљака и емитује хемијске сигнале за које се верује да помажу у комуникацији са мравима.[ 6]
Потпородице
Mating Cyaniris semiargus
Гусеница атале (Eumaeus atala )
Плавњак (Amblypodia anita ) из Theclinae
Највећа потпородица плаваца у Европи је потпородица правих плаваца (Polyommatinae ), са 113 врста у Европи. Затим следе репкари (Theclinae ) са 40 врста у Европи, дукати (Lycaeninae ) са 16 врста у Европи и Aphnaeinae , са једном врстом у Европи. У Србији живи 38 врста из подпородице Polyommatinae, 9 врста из подпородице Lycaeninae и 8 врста из подпородице Theclinae.
Многи таксономисти укључују само Lycaeninae , Theclinae , Polyommatinae , Poritiinae , Miletinae , и Curetinae под Lycaenidae.[ 7] [ 8] Aphnaeinae , који су некада били племе (Aphnaeini) унутар Theclinae, недавно су такође добили ранг подпородице.[ 9]
Неке старије класификације су укључивале и друге потпородице као што су Liphyrinae (сада Liphyrini, племе унутар Miletinae ), Lipteninae (сада Liptenini, племе унутар Poritiinae ) или Riodininae (сада посебна породица: Riodinidae ).
Овде се обично (али не и једнозначно) смешта фосилни род Lithodryas ; Lithopsyche је понекад смештен овде, али понекад у Riodininae.
Врсте плаваца
Најчешћи плавац, како у Европи, тако и у Србији је обични плавац (Polyommatus icarus ), остале честе врсте су стооки плавац (Plebejus argus ), мали дукат (Lycaena phlaeas ) и развигор (Aricia agestis ). Неки од најређих врста у Србији су: пакленац (Lycaena helle ), мочварни мравник (Phengaris teleius ), махунар (Iolana iolas ) и боровничар (Agriades optilete ). Најмањи плавац у Европи је Chilades trochylus . Последње откривени плавац у Србији је врста Драчац - Tarucus balkanicus. Пронађен је на крајњем југу Србије у границама заштићеног подручја Предео изузетних одлика "Долина Пчиње" 2021.године.[ 15] [ 16]
Allosmaitia strophius (Godart, 1824)
Arawacus jada (Hewitson, 1867)
Atlides halesus (Cramer, 1777)
Brephidium exilis (Boisduval, 1852)
Brephidium pseudofea (Morrison, 1873)
Callophrys affinis (W. H. Edwards, 1862)
Callophrys augustinus (Westwood, 1852)
Callophrys dumetorum (Boisduval, 1852)
Callophrys eryphon (Boisduval, 1852)
Callophrys fotis (Strecker, 1878)
Callophrys gryneus (Hübner, 1819)
Callophrys henrici (Grote and Robinson, 1867)
Callophrys hesseli (Rawson and Ziegler, 1950)
Callophrys irus (Godart, 1824)
Callophrys johnsoni (Skinner, 1904)
Callophrys lanoraieensis (Sheppard, 1934)
Callophrys mcfarlandi P. Ehrlich and Clench, 1960
Callophrys mossii (Hy. Edwards, 1881)
Callophrys muiri (Hy. Edwards, 1881)
Callophrys niphon (Hübner, 1819)
Callophrys perplexa W. Barnes and Benjamin, 1923
Callophrys polios (Cook and F. Watson, 1907)
Callophrys sheridanii (W. H. Edwards, 1877)
Callophrys spinetorum (Hewitson, 1867)
Callophrys thornei (J. Brown, 1983)
Callophrys xami (Reakirt, 1867)
Calycopis cecrops (Fabricius, 1793)
Calycopis isobeon (Butler and H. Druce, 1872)
Celastrina argiolus (Linnaeus, 1758)
Celastrina echo (W. H. Edwards, 1864)
Celastrina humulus Scott and D. Wright, 1998
Celastrina idella D. Wright and Pavulaan, 1999
Celastrina ladon (Cramer, 1780)
Celastrina lucia (W. Kirby, 1837)
Celastrina neglecta (W. H. Edwards, 1862)
Celastrina neglectamajor Opler and Krizek, 1984
Celastrina nigra (W. Forbes, 1960)
Celastrina serotina Pavulaan and D. Wright, 2005
Chlorostrymon maesites (Herrich-Schäffer, 1864)
Chlorostrymon simaethis (Drury, 1773)
Chlorostrymon telea (Hewitson, 1868)
Cupido amyntula (Boisduval, 1852)
Cupido comyntas (Godart, 1824)
Cyanophrys amyntor (Cramer, 1775)
Cyanophrys goodsoni (Clench, 1946)
Cyanophrys herodotus (Fabricius, 1793)
Cyanophrys miserabilis (Clench, 1946)
Cyclargus ammon (Lucas, 1857)
Cyclargus thomasi (Clench, 1941)
Echinargus isola (Reakirt, 1867)
Electrostrymon angelia (Hewitson, 1874)
Electrostrymon hugon (Godart, 1824)
Electrostrymon joya (Dognin, 1895)
Erora laeta (W. H. Edwards, 1862)
Erora quaderna (Hewitson, 1868)
Eumaeus atala (Poey, 1832)
Eumaeus toxea (Godart, 1824)
Euphilotes ancilla (W. Barnes and McDunnough, 1918)
Euphilotes battoides (Behr, 1867)
Euphilotes baueri (Shields, 1975)
Euphilotes bernardino (W. Barnes and McDunnough, 1916)
Euphilotes centralis (W. Barnes and McDunnough, 1917)
Euphilotes columbiae (Mattoni, 1954)
Euphilotes ellisii (Shields, 1975)
Euphilotes enoptes (Boisduval, 1852)
Euphilotes glaucon (W. H. Edwards, 1871)
Euphilotes mojave (F. Watson and W. Comstock, 1920)
Euphilotes pallescens (Tilden and Downey, 1955)
Euphilotes rita (W. Barnes and McDunnough, 1916)
Euphilotes spaldingi (W. Barnes and McDunnough, 1917)
Feniseca tarquinius (Fabricius, 1793)
Glaucopsyche lygdamus (E. Doubleday, 1841)
Glaucopsyche piasus (Boisduval, 1852)
Habrodais grunus (Boisduval, 1852)
Hemiargus ceraunus (Fabricius, 1793)
Hypaurotis crysalus (W. H. Edwards, 1873)
Hypostrymon critola (Hewitson, 1874)
Lampides boeticus (Linnaeus, 1767)
Leptotes cassius (Cramer, 1775)
Leptotes marina (Reakirt, 1868)
Lycaena arota (Boisduval, 1852)
Lycaena cupreus (W. H. Edwards, 1870)
Lycaena dione (Scudder, 1868)
Lycaena dorcas W. Kirby, 1837
Lycaena dospassosi McDunnough, 1940
Lycaena editha (Mead, 1878)
Lycaena epixanthe (Boisduval and Le Conte, 1835)
Lycaena gorgon (Boisduval, 1852)
Lycaena helloides (Boisduval, 1852)
Lycaena hermes (W. H. Edwards, 1870)
Lycaena heteronea Boisduval, 1852
Lycaena hyllus (Cramer, 1775)
Lycaena mariposa (Reakirt, 1866)
Lycaena nivalis (Boisduval, 1869)
Lycaena phlaeas (Linnaeus, 1761)
Lycaena rubidus (Behr, 1866)
Lycaena xanthoides (Boisduval, 1852)
Ministrymon azia (Hewitson, 1873)
Ministrymon clytie (W. H. Edwards, 1877)
Ministrymon leda (W. H. Edwards, 1882)
Ocaria ocrisia (Hewitson, 1868)
Oenomaus ortygnus (Cramer, 1779)
Parrhasius m-album (Boisduval and Le Conte, 1833)
Phaeostrymon alcestis (W. H. Edwards, 1871)
Philotes sonorensis (C. Felder and R. Felder, 1865)
Philotiella leona Hammond and McCorkle, 2000
Philotiella speciosa (Hy. Edwards, 1877)
Plebejus acmon (Westwood, 1851)
Plebejus anna (W. H. Edwards, 1861)
Plebejus argyrognomon (Bergsträsser, 1779)
Plebejus emigdionis (F. Grinnell, 1905)
Plebejus fridayi F. Chermock, 1945
Plebejus glandon (de Prunner, 1798)
Plebejus icarioides (Boisduval, 1852)
Plebejus idas (Linnaeus, 1761)
Plebejus lupini (Boisduval, 1869)
Plebejus melissa (W. H. Edwards, 1873)
Plebejus neurona (Skinner, 1902)
Plebejus optilete (Knoch, 1781)
Plebejus podarce (C. Felder and R. Felder, 1865)
Plebejus saepiolus (Boisduval, 1852)
Plebejus shasta (W. H. Edwards, 1862)
Rekoa marius (Lucas, 1857)
Rekoa palegon (Cramer, 1780)
Satyrium acadica (W. H. Edwards, 1862)
Satyrium auretorum (Boisduval, 1852)
Satyrium behrii (W. H. Edwards, 1870)
Satyrium calanus (Hübner, 1809)
Satyrium californica (W. H. Edwards, 1862)
Satyrium caryaevorus (McDunnough, 1942)
Satyrium edwardsii (Grote and Robinson, 1867)
Satyrium favonius (J. E. Smith, 1797)
Satyrium fuliginosa (W. H. Edwards, 1861)
Satyrium ilavia (Beutenmüller, 1899)
Satyrium kingi (Klots and Clench, 1952)
Satyrium liparops (Le Conte, 1833)
Satyrium polingi (W. Barnes and Benjamin, 1926)
Satyrium saepium (Boisduval, 1852)
Satyrium semiluna Klots, 1930
Satyrium sylvinus (Boisduval, 1852)
Satyrium tetra (W. H. Edwards, 1870)
Satyrium titus (Fabricius, 1793)
Strephonota tephraeus (Geyer, 1837)
Strymon acis (Drury, 1773)
Strymon albata (C. Felder and R. Felder, 1865)
Strymon alea (Godman and Salvin, 1887)
Strymon avalona (W. G. Wright, 1905)
Strymon bazochii (Godart, 1824)
Strymon bebrycia (Hewitson, 1868)
Strymon cestri (Reakirt, 1867)
Strymon istapa (Reakirt, 1867)
Strymon limenia (Hewitson, 1868)
Strymon martialis (Herrich-Schäffer, 1864)
Strymon melinus Hübner, 1818
Strymon rufofusca (Hewitson, 1877)
Strymon serapio (Godman and Salvin, 1887)
Strymon yojoa (Reakirt, 1867)
Tmolus echion (Linnaeus, 1767)
Udara blackburni (Tuely, 1878)
Ziegleria guzanta (Schaus, 1902)
Zizula cyna (W. H. Edwards, 1881)
Подврсте
Atlides halesus corcorani Clench, 1942
Atlides halesus halesus (Cramer, 1777)
Brephidium exilis exilis (Boisduval, 1852)
Brephidium exilis isophthalma (Herrich-Schäffer, 1862)
Brephidium pseudofea insularis Pavulaan and Gatrelle, 1999
Brephidium pseudofea pseudofea (Morrison, 1873)
Callophrys affinis affinis (W. H. Edwards, 1862)
Callophrys affinis albipalpis Gorelick, 2005
Callophrys affinis apama (W. H. Edwards, 1882)
Callophrys affinis homoperplexa W. Barnes and Benjamin, 1923
Callophrys affinis washingtonia Clench, 1944
Callophrys augustinus annetteae (dos Passos, 1943)
Callophrys augustinus augustinus (Westwood, 1852)
Callophrys augustinus concava (Austin in T. Emmel, 1998)
Callophrys augustinus croesioides (Scudder, 1876)
Callophrys augustinus helenae (dos Passos, 1943)
Callophrys augustinus iroides (Boisduval, 1852)
Callophrys dumetorum dumetorum (Boisduval, 1852)
Callophrys eryphon eryphon (Boisduval, 1852)
Callophrys eryphon fusca (Austin in T. Emmel, 1998)
Callophrys eryphon pallescens (Austin in T. Emmel, 1998)
Callophrys eryphon purpurascens (Austin and J. Emmel in T. Emmel, 1998)
Callophrys eryphon sheltonensis (F. Chermock and Frechin, 1949)
Callophrys fotis fotis (Strecker, 1878)
Callophrys fotis mojavensis (Austin in T. Emmel, 1998)
Callophrys gryneus byrnei K. Johnson, 1976
Callophrys gryneus castalis (W. H. Edwards, 1871)
Callophrys gryneus chalcosiva Clench, 1981
Callophrys gryneus gryneus (Hübner, 1819)
Callophrys gryneus juniperaria (J. A. Comstock, 1925)
Callophrys gryneus loki (Skinner, 1907)
Callophrys gryneus mansfieldi (Tilden, 1951)
Callophrys gryneus nelsoni (Boisduval, 1869)
Callophrys gryneus plicataria K. Johnson, 1976
Callophrys gryneus rosneri K. Johnson, 1976
Callophrys gryneus siva (W. H. Edwards, 1874)
Callophrys gryneus smilacis (Boisduval and Le Conte, 1835)
Callophrys gryneus sweadneri (F. Chermock, 1945)
Callophrys henrici henrici (Grote and Robinson, 1867)
Callophrys henrici margaretae (dos Passos, 1943)
Callophrys henrici solatus (Cook and F. Watson, 1909)
Callophrys henrici turneri (Clench, 1943)
Callophrys henrici viridissima (Pavulaan, 1998)
Callophrys henrici yahwehus (Gatrelle, 1999)
Callophrys hesseli angulata (Gatrelle, 2001)
Callophrys hesseli hesseli (Rawson and Ziegler, 1950)
Callophrys irus arsace (Boisduval and Le Conte, 1835)
Callophrys irus hadros (Cook and F. Watson, 1907)
Callophrys irus irus (Godart, 1824)
Callophrys mossii bayensis R. Brown, 1969
Callophrys mossii doudoroffi (dos Passos, 1940)
Callophrys mossii hidakupa (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Callophrys mossii marinensis (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Callophrys mossii mossii (Hy. Edwards, 1881)
Callophrys mossii schryveri (Cross, 1937)
Callophrys mossii windi (Clench, 1943)
Callophrys niphon clarki (T. Freeman, 1938)
Callophrys niphon niphon (Hübner, 1819)
Callophrys perplexa oregonensis Gorelick, 1970
Callophrys perplexa perplexa W. Barnes and Benjamin, 1923
Callophrys perplexa superperplexa J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Callophrys polios maritima (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Callophrys polios obscurus (Ferris and M. Fisher, 1973)
Callophrys polios polios (Cook and Watson, 1907)
Callophrys sheridanii comstocki Henne, 1940
Callophrys sheridanii interrupta Austin in T. Emmel, 1998
Callophrys sheridanii lemberti Tilden, 1963
Callophrys sheridanii neoperplexa W. Barnes and Benjamin, 1923
Callophrys sheridanii newcomeri Clench, 1963
Callophrys sheridanii paradoxa Scott, 1986
Callophrys sheridanii pseudodumetorum J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Callophrys sheridanii sacramento Scott, 2006
Callophrys sheridanii sheridanii (W. H. Edwards, 1877)
Callophrys spinetorum millerorum Clench, 1981
Callophrys spinetorum spinetorum (Hewitson, 1867)
Callophrys xami texami Clench, 1981
Callophrys xami xami (Reakirt, 1867)
Celastrina argiolus argiolus (Linnaeus, 1758)
Celastrina echo cinerea (W. H. Edwards, 1883)
Celastrina echo echo (W. H. Edwards, 1864)
Celastrina echo nigrescens (J. Fletcher, 1903)
Celastrina echo sidara (Clench, 1944)
Celastrina lucia lucia (W. Kirby, 1837)
Celastrina lucia lumarco Scott, 2006
Chlorostrymon simaethis sarita (Skinner, 1895)
Chlorostrymon simaethis simaethis (Drury, 1773)
Cupido amyntula albrighti (Clench, 1944)
Cupido amyntula amyntula (Boisduval, 1852)
Cupido amyntula herri (F. Grinnell, 1901)
Cupido amyntula maritima (Leblanc, 1985)
Cupido amyntula montanorum (Austin in T. Emmel, 1998)
Cupido amyntula nesiotes (J. Emmel and T. Emmel in T. Emmel, 1998)
Cupido amyntula valeriae (Clench, 1944)
Cupido comyntas comyntas (Godart, 1824)
Cupido comyntas sissona (W. G. Wright, 1905)
Cupido comyntas texana (F. Chermock, 1945)
Cyclargus thomasi bethunebakeri (W. Comstock and Huntington, 1943)
Electrostrymon angelia angelia (Hewitson, 1874)
Erora quaderna sanfordi dos Passos, 1940
Euphilotes ancilla ancilla (W. Barnes and McDunnough, 1918)
Euphilotes ancilla gilvatunica Austin in T. Emmel, 1998
Euphilotes ancilla giulianii Pratt and J. Emmel in T. Emmel, 1998
Euphilotes ancilla pseudointermedia Pratt and J. Emmel in T. Emmel, 1998
Euphilotes ancilla purpura Austin in T. Emmel, 1998
Euphilotes ancilla shieldsi Austin in T. Emmel, 1998
Euphilotes battoides allyni (Shields, 1975)
Euphilotes battoides anasazi Scott, 1998
Euphilotes battoides argocyanea Pratt and J. Emmel in T. Emmel, 1998
Euphilotes battoides battoides (Behr, 1867)
Euphilotes battoides fusimaculata Austin in T. Emmel, 1998
Euphilotes battoides mazourka Pratt and J. Emmel in T. Emmel, 1998
Euphilotes battoides panamintensis Pratt and J. Emmel in T. Emmel, 1998
Euphilotes battoides vernalis Pratt and J. Emmel in T. Emmel, 1998
Euphilotes baueri baueri (Shields, 1975)
Euphilotes baueri orientis Austin in T. Emmel, 1998
Euphilotes bernardino bernardino (W. Barnes and McDunnough, 1916)
Euphilotes bernardino inyomontana Pratt and J. Emmel in T. Emmel, 1998
Euphilotes bernardino martini (Mattoni, 1954)
Euphilotes bernardino minuta Austin in T. Emmel, 1998
Euphilotes centralis centralis (W. Barnes and McDunnough, 1917)
Euphilotes centralis hadrochilus Pratt and J. Emmel in T. Emmel, 1998
Euphilotes ellisii avawatz Pratt and J. Emmel in T. Emmel, 1998
Euphilotes ellisii basinensis Austin in T. Emmel, 1998
Euphilotes ellisii ellisii (Shields, 1975)
Euphilotes ellisii euromojavensis Pratt and J. Emmel in T. Emmel, 1998
Euphilotes enoptes arenacola Pratt and J. Emmel in T. Emmel, 1998
Euphilotes enoptes aridorum Austin in T. Emmel, 1998
Euphilotes enoptes bayensis (Langston, 1964)
Euphilotes enoptes cryptorufes Pratt and J. Emmel in T. Emmel, 1998
Euphilotes enoptes dammersi (J. A. Comstock and Henne, 1933)
Euphilotes enoptes enoptes (Boisduval, 1852)
Euphilotes enoptes langstoni (Shields, 1975)
Euphilotes enoptes opacapulla Austin in T. Emmel, 1998
Euphilotes enoptes primavera Austin in T. Emmel, 1998
Euphilotes enoptes smithi (Mattoni, 1954)
Euphilotes enoptes tildeni (Langston, 1964)
Euphilotes glaucon australoglaucon Pratt and J. Emmel in T. Emmel, 1998
Euphilotes glaucon comstocki (Shields, 1975)
Euphilotes glaucon glaucon (W. H. Edwards, 1871)
Euphilotes glaucon intermedia (W. Barnes and McDunnough, 1917)
Euphilotes glaucon oregonensis (W. Barnes and McDunnough, 1917)
Euphilotes mojave mojave (F. Watson and W. Comstock, 1920)
Euphilotes mojave virginensis Austin in T. Emmel, 1998
Euphilotes pallescens arenamontana Austin in T. Emmel, 1998
Euphilotes pallescens calneva J. Emmel and T. Emmel in T. Emmel, 1998
Euphilotes pallescens confusa Pratt and J. Emmel in T. Emmel, 1998
Euphilotes pallescens elvirae (Mattoni, 1966)
Euphilotes pallescens emmeli (Shields, 1975)
Euphilotes pallescens mattonii (Shields, 1975)
Euphilotes pallescens pallescens (Tilden and Downey, 1955)
Euphilotes pallescens ricei Austin in T. Emmel, 1998
Euphilotes rita coloradensis (Mattoni, 1966)
Euphilotes rita rita (W. Barnes and McDunnough, 1916)
Euphilotes spaldingi pinjuna Scott, 1981
Euphilotes spaldingi spaldingi (W. Barnes and McDunnough, 1917)
Feniseca tarquinius novascotiae McDunnough, 1935
Feniseca tarquinius tarquinius (Fabricius, 1793)
Glaucopsyche lygdamus afra (W. H. Edwards, 1884)
Glaucopsyche lygdamus arizonensis McDunnough, 1936
Glaucopsyche lygdamus australis F. Grinnell, 1917
Glaucopsyche lygdamus columbia (Skinner, 1917)
Glaucopsyche lygdamus couperi Grote, 1873
Glaucopsyche lygdamus deserticola Austin and J. Emmel in T. Emmel, 1998
Glaucopsyche lygdamus incognitus Tilden, 1974
Glaucopsyche lygdamus jacki D. Stallings and Turner, 1947
Glaucopsyche lygdamus lygdamus (E. Doubleday, 1841)
Glaucopsyche lygdamus mildredae F. Chermock, 1945
Glaucopsyche lygdamus minipunctum Austin in T. Emmel, 1998
Glaucopsyche lygdamus orcus (W. H. Edwards, 1869)
Glaucopsyche lygdamus oro (Scudder, 1876)
Glaucopsyche lygdamus palosverdesensis E. Perkins and J. Emmel, 1977
Glaucopsyche lygdamus pseudoxerces J. Emmel and T. Emmel in T. Emmel, 1998
Glaucopsyche lygdamus sabulosa J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Glaucopsyche lygdamus xerces (Boisduval, 1852)
Glaucopsyche piasus daunia (W. H. Edwards, 1871)
Glaucopsyche piasus excubita J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Glaucopsyche piasus gabrielina J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Glaucopsyche piasus nevada F. Brown, 1975
Glaucopsyche piasus piasus (Boisduval, 1852)
Glaucopsyche piasus sagittigera (C. Felder and R. Felder, 1865)
Glaucopsyche piasus toxeuma F. Brown, 1971
Glaucopsyche piasus umbrosa J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Habrodais grunus grunus (Boisduval, 1852)
Habrodais grunus herri W. D. Field, 1938
Habrodais grunus lorquini W. D. Field, 1938
Hemiargus ceraunus antibubastus Hübner, 1818
Hemiargus ceraunus astenidas (Lucas, 1857)
Hemiargus ceraunus gyas (W. H. Edwards, 1871)
Hypaurotis crysalus citima (Hy. Edwards, 1881)
Hypaurotis crysalus crysalus (W. H. Edwards, 1873)
Hypaurotis crysalus intermedia Austin in T. Emmel, 1998
Leptotes cassius cassidula (Boisduval, 1870)
Leptotes cassius theonus (Lucas, 1857)
Lycaena arota arota (Boisduval, 1852)
Lycaena arota nubila (J. A. Comstock, 1926)
Lycaena arota schellbachi (Tilden, 1955)
Lycaena arota virginiensis (W. H. Edwards, 1870)
Lycaena cupreus cupreus (W. H. Edwards, 1870)
Lycaena cupreus lapidicola J. Emmel and Pratt in T. Emmel, 1998
Lycaena cupreus snowi (W. H. Edwards, 1881)
Lycaena dorcas arcticus (Ferris, 1977)
Lycaena dorcas castro (Reakirt, 1866)
Lycaena dorcas claytoni A. E. Brower, 1940
Lycaena dorcas dorcas W. Kirby, 1837
Lycaena dorcas florus (W. H. Edwards, 1884)
Lycaena dorcas megaloceras (Ferris, 1977)
Lycaena dorcas michuron Scott, 2006
Lycaena editha editha (Mead, 1878)
Lycaena editha obscuramaculata Austin, 1989
Lycaena editha pseudonexa J. Emmel and Pratt in T. Emmel, 1998
Lycaena editha vurali Koçak, 1984
Lycaena epixanthe epixanthe (Boisduval and Le Conte, 1835)
Lycaena epixanthe michiganensis Rawson, 1948
Lycaena epixanthe phaedrus (G. Hall, 1924)
Lycaena gorgon dorothea J. Emmel and Pratt in T. Emmel, 1998
Lycaena gorgon gorgon (Boisduval, 1852)
Lycaena gorgon jacquelineae J. Emmel and Pratt in T. Emmel, 1998
Lycaena gorgon micropunctata J. Emmel and Pratt in T. Emmel, 1998
Lycaena heteronea clara Hy. Edwards, 1877
Lycaena heteronea gravenotata Klots, 1930
Lycaena heteronea heteronea Boisduval, 1852
Lycaena heteronea klotsi W. D. Field, 1936
Lycaena heteronea northi J. Emmel and Pratt in T. Emmel, 1998
Lycaena heteronea rava Austin in T. Emmel, 1998
Lycaena heteronea rutila Austin in T. Emmel, 1998
Lycaena heteronea submaculata J. Emmel and Pratt in T. Emmel, 1998
Lycaena mariposa charlottensis (W. Holland, 1930)
Lycaena mariposa mariposa (Reakirt, 1866)
Lycaena mariposa penroseae W. D. Field, 1938
Lycaena nivalis bichroma J. Emmel and Pratt in T. Emmel, 1998
Lycaena nivalis browni dos Passos, 1938
Lycaena nivalis nivalis (Boisduval, 1869)
Lycaena nivalis praetexta Austin in T. Emmel, 1998
Lycaena nivalis warnermontana J. Emmel and Pratt in T. Emmel, 1998
Lycaena phlaeas alpestris J. Emmel and Pratt in T. Emmel, 1998
Lycaena phlaeas arctodon Ferris, 1974
Lycaena phlaeas arethusa (Dod, 1907)
Lycaena phlaeas feildeni (M'Lachlan, 1878)
Lycaena phlaeas hypophlaeas (Boisduval, 1852)
Lycaena rubidus duofascies K. Johnson and Balogh, 1977
Lycaena rubidus ferrisi K. Johnson and Balogh, 1977
Lycaena rubidus incana Austin in T. Emmel, 1998
Lycaena rubidus longi K. Johnson and Balogh, 1977
Lycaena rubidus monachensis K. Johnson and Balogh, 1977
Lycaena rubidus perkinsorum K. Johnson and Balogh, 1977
Lycaena rubidus rubidus (Behr, 1866)
Lycaena rubidus sirius (W. H. Edwards, 1871)
Lycaena xanthoides nigromaculata J. Emmel and Pratt in T. Emmel, 1998
Lycaena xanthoides obsolescens J. Emmel and Pratt in T. Emmel, 1998
Lycaena xanthoides xanthoides (Boisduval, 1852)
Phaeostrymon alcestis alcestis (W. H. Edwards, 1871)
Phaeostrymon alcestis oslari (Dyar, 1904)
Philotes sonorensis extinctis Mattoni, 1991
Philotes sonorensis sonorensis (C. Felder and R. Felder, 1865)
Philotiella speciosa bohartorum (Tilden, 1969)
Philotiella speciosa purisima Priestaf and J. Emmel in T. Emmel, 1998
Philotiella speciosa septentrionalis Austin in T. Emmel, 1998
Philotiella speciosa speciosa (Hy. Edwards, 1877)
Plebejus anna anna (W. H. Edwards, 1861)
Plebejus anna azureus (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus anna benwarner Scott, 2006
Plebejus anna lotis (Lintner, 1878)
Plebejus anna ricei Cross, 1937
Plebejus anna vancouverensis (C. Guppy and J. Shepard, 2001)
Plebejus argyrognomon argyrognomon (Bergsträsser, 1779)
Plebejus glandon bryanti Leussler, 1935
Plebejus glandon cassiope (J. Emmel and T. Emmel in T. Emmel, 1998)
Plebejus glandon franklinii (J. Curtis, 1835)
Plebejus glandon kelsoni (J. Emmel and T. Emmel in T. Emmel, 1998)
Plebejus glandon labrador (Schmidt, Scott and Kondla, 2006)
Plebejus glandon lacustris T. Freeman, 1939
Plebejus glandon megalo McDunnough, 1927
Plebejus glandon punctatus (Austin in T. Emmel, 1998)
Plebejus glandon rustica (W. H. Edwards, 1865)
Plebejus icarioides albihalos (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus icarioides argusmontana (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus icarioides atascadero (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus icarioides austinorum (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus icarioides blackmorei W. Barnes and McDunnough, 1919
Plebejus icarioides buchholzi dos Passos, 1938
Plebejus icarioides eosierra (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus icarioides evius (Boisduval, 1869)
Plebejus icarioides fenderi Macy, 1931
Plebejus icarioides fulla (W. H. Edwards, 1870)
Plebejus icarioides helios (W. H. Edwards, 1871)
Plebejus icarioides icarioides (Boisduval, 1852)
Plebejus icarioides inyo (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus icarioides lycea (W. H. Edwards, 1864)
Plebejus icarioides missionensis Hovanitz, 1937
Plebejus icarioides montis Blackmore, 1923
Plebejus icarioides moroensis Sternitzky, 1930
Plebejus icarioides panamintina (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus icarioides parapheres (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus icarioides pardalis (Behr, 1867)
Plebejus icarioides pembina (W. H. Edwards, 1862)
Plebejus icarioides pheres (Boisduval, 1852)
Plebejus icarioides santana (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus idas alaskensis F. Chermock, 1945
Plebejus idas aster (W. H. Edwards, 1882)
Plebejus idas atrapraetextus W. D. Field, 1939
Plebejus idas empetri T. Freeman, 1938
Plebejus idas ferniensis F. Chermock, 1945
Plebejus idas longinus (Nabokov, 1949)
Plebejus idas nabokovi (Masters, 1972)
Plebejus idas scudderii (W. H. Edwards, 1861)
Plebejus idas sublivens (Nabokov, 1949)
Plebejus lupini alpicola (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus lupini argentata (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus lupini chlorina (Skinner, 1902)
Plebejus lupini cotundra Scott, 2006
Plebejus lupini dedeckera (J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998)
Plebejus lupini goodpasturei (Austin in T. Emmel, 1998)
Plebejus lupini lupini (Boisduval, 1869)
Plebejus lupini lutzi dos Passos, 1938
Plebejus lupini monticola (Clemence, 1909)
Plebejus lupini spangelatus Burdick, 1942
Plebejus lupini texanus (Goodpasture, 1973)
Plebejus melissa alateres (Austin in T. Emmel, 1998)
Plebejus melissa annetta (W. H. Edwards, 1882)
Plebejus melissa inyoensis (Nabokov, 1949)
Plebejus melissa melissa (W. H. Edwards, 1873)
Plebejus melissa paradoxa F. Chermock, 1945
Plebejus melissa pseudosamuelis (Nabokov, 1949)
Plebejus melissa samuelis (Nabokov, 1944)
Plebejus optilete yukona (W. Holland, 1900)
Plebejus podarce cilla (Behr, 1867)
Plebejus podarce klamathensis (J. Emmel and T. Emmel in T. Emmel, 1998)
Plebejus podarce podarce (C. Felder and R. Felder, 1865)
Plebejus saepiolus aehaja (Behr, 1867)
Plebejus saepiolus albomontanus Austin and J. Emmel in T. Emmel, 1998
Plebejus saepiolus amica (W. H. Edwards, 1863)
Plebejus saepiolus aureolus J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Plebejus saepiolus gertschi dos Passos, 1938
Plebejus saepiolus hilda (J. Grinnell and F. Grinnell, 1907)
Plebejus saepiolus insulanus Blackmore, 1920
Plebejus saepiolus littoralis J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Plebejus saepiolus maculosus Austin in T. Emmel, 1998
Plebejus saepiolus rufescens (Boisduval, 1869)
Plebejus saepiolus saepiolus (Boisduval, 1852)
Plebejus saepiolus whitmeri F. Brown, 1951
Plebejus shasta calchas (Behr, 1867)
Plebejus shasta charlestonensis Austin, 1980
Plebejus shasta minnehaha (Scudder, 1874)
Plebejus shasta pallidissima (Austin in T. Emmel, 1998)
Plebejus shasta pitkinensis Ferris, 1976
Plebejus shasta platazul Scott, 2006
Plebejus shasta shasta (W. H. Edwards, 1862)
Satyrium acadica acadica (W. H. Edwards, 1862)
Satyrium acadica coolinensis (F. Watson and W. Comstock, 1920)
Satyrium acadica montanensis (F. Watson and W. Comstock, 1920)
Satyrium auretorum auretorum (Boisduval, 1852)
Satyrium auretorum fumosum J. Emmel and Mattoni, 1990
Satyrium auretorum spadix (Hy. Edwards, 1881)
Satyrium behrii behrii (W. H. Edwards, 1870)
Satyrium behrii columbia (McDunnough, 1944)
Satyrium behrii crossi (W. D. Field, 1938)
Satyrium calanus albidus Scott, 1981
Satyrium calanus calanus (Hübner, 1809)
Satyrium calanus falacer (Godart, 1824)
Satyrium calanus godarti (W. D. Field, 1938)
Satyrium californica brashor Kondla and Scott, 2006
Satyrium californica californica (W. H. Edwards, 1862)
Satyrium californica cygnus (W. H. Edwards, 1871)
Satyrium californica obscurafacies Austin in T. Emmel, 1998
Satyrium californica wapiti M. Fisher, 2006
Satyrium edwardsii edwardsii (Grote and Robinson, 1867)
Satyrium edwardsii meridionale Gatrelle, 2001
Satyrium favonius autolycus (W. H. Edwards, 1871)
Satyrium favonius favonius (J. E. Smith, 1797)
Satyrium favonius ontario (W. H. Edwards, 1868)
Satyrium favonius violae (D. Stallings and Turner, 1947)
Satyrium fuliginosa albolineatum Mattoon and Austin in T. Emmel, 1998
Satyrium fuliginosa fuliginosa (W. H. Edwards, 1861)
Satyrium fuliginosa tildeni Mattoon and Austin in T. Emmel, 1998
Satyrium liparops aliparops (Michener and dos Passos, 1942)
Satyrium liparops fletcheri (Michener and dos Passos, 1942)
Satyrium liparops floridensis Gatrelle, 2001
Satyrium liparops liparops (Le Conte, 1833)
Satyrium liparops strigosa (T. Harris, 1862)
Satyrium polingi organensis Ferris, 1980
Satyrium polingi polingi (W. Barnes and Benjamin, 1926)
Satyrium saepium caliginosum J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Satyrium saepium chalcis (W. H. Edwards, 1869)
Satyrium saepium chlorophora (Watson and W. Comstock, 1920)
Satyrium saepium fulvescens (Hy. Edwards, 1877)
Satyrium saepium latalinea Austin and Savage, 1998
Satyrium saepium obscurofuscum Austin in T. Emmel, 1998
Satyrium saepium provo (F. Watson and W. Comstock, 1920)
Satyrium saepium rubrotenebrosum J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Satyrium saepium saepium (Boisduval, 1852)
Satyrium saepium subaridum J. Emmel, T. Emmel and Mattoon in T. Emmel, 1998
Satyrium semiluna maculadistinctum Mattoon and Austin in T. Emmel, 1998
Satyrium semiluna semiluna Klots, 1930
Satyrium sylvinus desertorum (F. Grinnell, 1917)
Satyrium sylvinus dryope (W. H. Edwards, 1870)
Satyrium sylvinus itys (W. H. Edwards, 1882)
Satyrium sylvinus megapallidum Austin in T. Emmel, 1998
Satyrium sylvinus nootka M. Fisher, 1998
Satyrium sylvinus putnami (Hy. Edwards, 1877)
Satyrium sylvinus sylvinus (Boisduval, 1852)
Satyrium titus campus (Gatrelle, 2004)
Satyrium titus immaculosus (W. Comstock, 1913)
Satyrium titus occidentalis (Austin and J. Emmel in T. Emmel, 1998)
Satyrium titus titus (Fabricius, 1793)
Satyrium titus watsoni (W. Barnes and Benjamin, 1926)
Satyrium titus winteri (Gatrelle, 2004)
Strymon acis bartrami (W. Comstock and Huntington, 1943)
Strymon bazochii bazochii (Godart, 1824)
Strymon bazochii gundlachianus M. Bates, 1935
Strymon istapa clenchi Austin and J. Emmel in T. Emmel, 1998
Strymon istapa istapa (Reakirt, 1867)
Strymon istapa modesta (Maynard, 1873)
Strymon melinus atrofasciata McDunnough, 1921
Strymon melinus franki W. D. Field, 1938
Strymon melinus humuli (T. Harris, 1842)
Strymon melinus melinus Hübner, 1818
Strymon melinus pudica (Hy. Edwards, 1877)
Strymon melinus setonia McDunnough, 1927
Tmolus echion echiolus (Draudt, 1920)
Галерија
Смарагдни репкар (Callophrys rubi )
Наранџасти репкар (Thecla betulae )
Најмањи лептир Европе - Chilades trochylus
Гусеница плавца Eumaeus atala
Плавац Curetis thetis
Краткорепи селац (Leptotes pirithous )
Развигор (Aricia agestis )
Блистави плавац (Lysandra belargus)
Загасити репкар (Satyrium ilicis )
Референце
^ „Биолошка разноврсност Србије” . bioras.petnica.rs. Архивирано из оригинала 12. 02. 2015. г. Приступљено 12. 01. 2018 .
^ Pierce, Naomi E.; Braby, Michael F.; Heath, Alan; Lohman, David J.; Mathew, John; Rand, Douglas B.; Travassos, Mark A. (2002). „The Ecology and Evolution of Ant Association in the Lycaenidae (Lepidoptera)” . Annual Review of Entomology . 47 (1): 259—267. PMID 11729090 . doi :10.1146/annurev.ento.47.091201.145257 .
^ а б в Pierce, N. E.; Braby, M. F.; Heath, A.; Lohman, D. J.; Mathew, J.; Rand, D. B.; Travassos, M. A. (2002). „"The ecology and evolution of ant association in the Lycaenidae (Lepidoptera)" ” (PDF) . Annual Review of Entomology . 47 : 733—771. PMID 11729090 . doi :10.1146/annurev.ento.47.091201.145257 . . .
'^ DeVries, Philip J (1992). „Singing Caterpillars, Ants and Symbiosis”. Scientific American, 267' .
^ Robbins, Robert K. (1981). „The 'False Head' Hypothesis: Predation and Wing Pattern Variation of Lycaenid Butterflies” . American Naturalist . 118 (5): 770—775. S2CID 34146954 . doi :10.1086/283868 .
^ "Lycaenid Butterflies and Ants" . Australian Museum . Archived 18 November 2007.
^ Brower, Andrew V. Z. (2008). "Lycaenidae [Leach] 1815" . Version 25 April 2008 (under construction). The Tree of Life Web Project .
^ Ackery, P. R.; de Jong, R. & Vane-Wright, R. I. (1999). "The butterflies: Hedyloidea, Hesperioidea, and Papilionoidea". Pages 264-300 in: Lepidoptera: Moths and Butterflies. 1. Evolution, Systematics, and Biogeography. Handbook of Zoology Vol. IV, Part 35. N. P. Kristensen, ed. De Gruyter, Berlin and New York.
^ а б в Boyle, John H.; Kaliszewska, Zofia A.; Espeland, Marianne; Suderman, Tamara R.; Fleming, Jake; Heath, Alan; Pierce, Naomi E. (2015). „Phylogeny of the Aphnaeinae: Myrmecophilous African butterflies with carnivorous and herbivorous life histories”. Systematic Entomology . 40 (1): 169—182. Bibcode :2015SysEn..40..169B . S2CID 83052979 . doi :10.1111/syen.12098 .
^ а б R.K., Varshney; Smetacek, Peter (2015). A Synoptic Catalogue of the Butterflies of India . New Delhi: Butterfly Research Centre, Bhimtal & Indinov Publishing, New Delhi. стр. 89. ISBN 978-81-929826-4-9 . doi :10.13140/RG.2.1.3966.2164 .
^ Beccaloni, G.; Scoble, M.; Kitching, I.; Simonsen, T.; Robinson, G.; Pitkin, B.; Hine, A.; Lyal, C., ур. (2003). „Плавци ” . The Global Lepidoptera Names Index . Natural History Museum . Retrieved April 20, 2018.
^ Markku Savela's website on Lepidoptera Curetis [мртва веза ] at Markku Savela's website on Lepidoptera
^ Lohman, David J.; Samarita, Venancio U. (2009). „The biology of carnivorous butterfly larvae (Lepidoptera: Lycaenidae: Miletinae: Miletini) and their ant-tended hemipteran prey in Thailand and the Philippines”. Journal of Natural History . 43 (9–10): 569—581. Bibcode :2009JNatH..43..569L . doi :10.1080/00222930802610485 .
^ Bingham, C.T. (1907). The Fauna of British India, Including Ceylon and Burma . II (1st изд.). London: Taylor and Francis, Ltd.
^ Tot Ivan; Djurić Milan; Vukajlović Filip; Vujić Mihailo (2021-11-11). „LITTLE TIGER BLUE, TARUCUS BALKANICUS (FREYER, 1845) – A NEW BUTTERFLY SPECIES IN THE FAUNA OF SERBIA (LEPIDOPTERA: LYCAENIDAE)” . Acta Entomologica Serbica (на језику: енглески). 26 (2): 85—88. doi :10.5281/ZENODO.5675734 .
^ „Tarucus balkanicus - Dračac | Alciphron” . alciphron.habiprot.org.rs . Приступљено 2024-10-03 .
Литература
Bridges, Charles A. (1994). Catalogue of the Family-Group, Genus-Group and Species-Group Names of the Riodinidae & Lycaenidae (Lepidoptera) of the World . Urbana, Ill.
Eliot, J. N. (1973). „"The higher classification of the Lycaenidae (Lepidoptera): a tentative arrangement" ” . 28 : 371—505. . Bulletin of the British Museum (Natural History) (Entomology) . .
Glassberg, Jeffrey (2001). Butterflies Through Binoculars, The West .
Guppy, Crispin S. & Shepard, Jon H. Butterflies of British Columbia . 2001.
James, David G. and Nunnallee, David (2011). Life Histories of Cascadia Butterflies .
Pelham, Jonathan (2008). Catalogue of the Butterflies of the United States and Canada .
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