Шкорпије [ 1] скорпије ,[ 1] шкорпиони [ 2] лат. Scorpiones) спадају у најкрупније пауколике животиње , а вероватно најстарије (фосили потичу још из силура ) и прве сувоземне зглавкаре . Насељавају углавном суптропске и тропске пределе мада их има и у умереној зони. Преко дана мирују сакривене испод лишћа и камења, а ноћу се активирају у потрази за храном. Величина им се креће од 13 mm до 25 cm. Углавном настањују пустиње, али су се прилагодиле великом броју различитих станишта. Присутне су на свим континентима осим Антарктика . Ред укључује 22 породице са преко 2.500 описаних врста.
Отров шкорпија је неуротоксичан и код већине врста није смртоносан за човека. Серум који се даје против отрова шкорпија ефикасан је само до одређеног времена након убода. Међутим, отров појединих врста може угрозити живот човека:
Androctonus australis Тело шкорпија подељено је на два региона прозому и опистозому које су широко спојене.
Прозома карапаксом . На леђној страни налазе се очи: један пар средишњих и неколико пари латералних. На трбушној страни нема оклопа већ је она заштићена основама ногу.
На прозоми се налазе следећи екстремитети :
пар су хелицере смештене испред усног отвора , у облику клешта, сићушне и изграђене од три сегмента ;
пар су педипалпи , снажно развијени, клештолико завршени и главни знак распознавања скорпија; служе за хватање плена;
четири пара ногу за ходање од којих се свака састоји од 7 чланака. Опистозома
предњег, преабдомена (мезозома) кога гради 7 чланака;
задњег, постабдомена (метазома) који је изграђен од 5 чланака, на његовом крају се налази повијена бодља. Опистозомални сегменти су сви добро заштићени спољашњим скелетом који је изграђен од 4 плоче:
тергит на дорзалној (леђној) страни;стернит , на вентралној (трбушној) страни;плеуре постављене са стране и мембранозне. На опистозоми се налазе остаци екстремитета:
Када шкорпија хвата плен онда је њен постабдомен подигнут и повијен напред преко тела (на слици) тако да је бодља управљена ка жртви и спремна за напад. Бодља, којом се завршава опистозома се, према ембрионалном пореклу, сматра чланком. У њеној проширеној основи налази се парна отровна жлезда чији се отвор налази на врху бодље. Жлезда је обавијена мишићима чије контракције убризгавају отров у жртву.
Шкорпије се хране ситним зглавкарима које хватају клештоликим педипалпима, а убијају убодом бодље.
Женка са младунцима на леђима Варење усмрћеног плена почиње радом хелицера које мацерирају плен и излучивањем сокова из предњег црева на њега. Полусварену храну усисавају ждрелом из кога храна иде дање у једњак па у средње црево и жлезде за варење где се вари до краја, а затим и апсорбује. Несварени састојци хране се пребацују у задње црево и преко аналног отвора избацују у спољашњу средину.
Излучивање се обавља помоћу:
1. два пара Малпигијевих цевчица које се изливају између средњег и задњег црева;
2. пара коксалних жлезда које се изливају у базу трећег пара ногу за ходање.
Крвни систем је отвореног типа и састоји се од:
срца смештено у перикардијуму у преабдомену; крв из перикардијума доспева у срце преко отвора остија ;артерија које одводе крв из срца у све делове тела;синуса , у које се крв из артерија излива;венозних канала, којима се крв из синуса враћа у срце.Дисање се врши помоћу 4 пара листоликих плућа која су утопљена у крви вентралног синуса. У њему се крв оксидује, а затим враћа у срце.
Нервни систем се састоји од:
Чулни органи скорпија су:
тактилни у облику чулних длачица различитих облика;очи еверзног типа;
чешљолики остаци екстремитета на којима се налазе чулне ћелије чија функција није довољно проучена. Шкорпије су одвојених полова са слабо израженим полним диморфизмом који се углавном огледа у томе што мужјаци на оперкулуму имају кукице и по облику бодљи и педипалпи. Гонаде код мужјака и женки су врло сличне грађе, састоје се од групе цевчица од које полазе одводи који се изливају у генитални атријум .
Приликом копулације шкорпије показују врло специфично понашање познато као игра парења . Мужјаци полажу сперматозоиде на земљу, а женке их уносе у тело преко полног отвора. Развиће оплођених јаја одвија се у телу женке и траје од неколико месеци до годину дана, а некада и дуже. После завршеног развића рађа се од 6 до 90 младих дужине неколико милиметара који на леђима мајке бораве око недељу дана, односно, до првог пресвлачења .
E. mingrelicus Ред Scorpiones је подељен на седам фамилија са преко 100 родова и око 700 врста. Најпознатије су следеће фамилије и родови:
род Androctonus
род Buthus
род Mesobuthus M. gibbosus живи на просторима бивше Југославије; породица Chactidae , као и претходна насељава скоро све континенте; припадају јој родови: Euscorpius E. italicus , E. carpathicus и E. migrelicus живе у Србији;Chactus Scorpio Pandinus
Akravidae Levy, 2007 Archaeobuthidae Lourenço, 2001 Belisariidae Lourenço, 1998 Bothriuridae Simon, 1880 Buthidae C. L. Koch, 1837 Caraboctonidae Kraepelin, 1905 Chactidae Pocock, 1893 Chaerilidae Pocock, 1893 Chaerilobuthidae Lourenço and Beigel, 2011 Diplocentridae Karsch, 1880 Euscorpiidae Laurie, 1896 Hadruridae Stahnke, 1974 Hemiscorpiidae Pocock, 1893 Heteroscorpionidae Kraepelin, 1905 Hormuridae Laurie, 1896 Iuridae Thorell, 1876 Palaeoburmesebuthidae Lourenço, 2015 Palaeoeuscorpiidae Lourenço, 2003 Palaeotrilineatidae Lourenço, 2012 Protoischnuridae Carvalho and Lourenço, 2001 Pseudochactidae Gromov, 1998 Rugodentidae Bastawade, Sureshan and Radhakrishnan, 2005 Scorpionidae Latreille, 1802 Scorpiopidae Kraepelin, 1905 Sucinlourencoidae Rossi, 2015 Superstitioniidae Stahnke, 1940 Vaejovidae Thorell, 1876
Alacraninae Vignoli and Prendini, 2009 Archaeoscorpiopinae Lourenço, 2003 Bothriurinae Simon, 1880 Calchinae Birula, 1917 Chaerilinae Pocock, 1893 Electrochaerilinae Santiago-Blay, Fet, Soleglad and Anderson, 2004 Euscorpiinae Laurie, 1896 Hemiscorpiinae Pocock, 1893 Heterometrinae Simon, 1879 Hormurinae Laurie, 1896 Iurinae Thorell, 1876 Lisposominae Lawrence, 1928 Megacorminae Opistophthalminae Rossi, 2016 Pandininae Thorell, 1876 Pseudochactinae Gromov, 1998 Rugodentinae Bastawade, Sureshan and Radhakrishnan, 2005 Scorpioninae Latreille, 1802 Scorpiopinae Kraepelin, 1905 Smeringurinae Soleglad and Fet, 2008 Superstitioniinae Stahnke, 1940 Syntropinae Kraepelin, 1905 Troglokhammouaninae Prendini, Ehrenthal and Loria, 2021 Typhlochactinae Mitchell, 1971 Urodacinae Pocock, 1893 Vaejovinae Thorell, 1876 Vietbocapinae Lourenço, 2012
Aegaeobuthus Kovarík, 2019 Aemngvantom Prendini, Ehrenthal and Loria, 2021 Afroisometrus Kovarík, 1997 Afrolychas Kovarík, 2019 Akentrobuthus Lamoral, 1976 Akrav Levy, 2007 Alacran Francke, 1982 Alayotityus Armas, 1973 Alpiscorpius Gantenbein, Fet, Largiadèr and Scholl, 1999 Ananteris Thorell, 1891 Ananteroides Borelli, 1911 Anatoliurus Parmakelis et al., 2022 Androctonus Ehrenberg, 1828 Anomalobuthus Kraepelin, 1900 Antridiscalceatus Rossi, 2018 Anuroctonus Pocock, 1893 Aops Volschenk and Prendini, 2008 Apistobuthus Finnegan, 1932 Archaeoananteroides Lourenço in Lourenço and Velten, 2016 Archaeobuthus Lourenço, 2001 Archaeoscorpiops Lourenço, 2015 Australobuthus Locket, 1990 Auyantepuia González-Sponga, 1978 Babycurus Karsch, 1886 Baloorthochirus Kovarík, 1996 Balsateres González-Santillán and Prendini, 2013 Barbaracurus Kovarík, Lowe and Stáhlavsky, 2018 Belisarius Simon, 1879 Betaburmesebuthus Lourenço in Lourenço and Beigel, 2015 Bioculus Stahnke, 1968 Birulatus Vachon, 1974 Bothriurus Simon, 1880 Brachistosternus Pocock, 1893 Brazilobothriurus Lourenço and Monod, 2000 Broteochactas Pocock, 1893 Brotheas C. L. Koch, 1837 Burmesescorpiops Lourenço, 2016 Buthacus Birula, 1908 Butheoloides Hirst, 1925 Butheolus Simon, 1882 Buthiscus Birula, 1905 Buthoscorpio Werner, 1936 Buthus Leach, 1815 Calchas Birula, 1899 Caraboctonus Pocock, 1893 Catalinia Soleglad, Ayrey, Graham and Fet, 2017 Cazierius Francke, 1978 Centromachetes Lönnberg, 1897 Centruroides Marx, 1890 Cercophonius Peters, 1861 Chactas Gervais, 1844 Chactopsis Kraepelin, 1912 Chactopsoides Ochoa, Rojas-Runjaic, Pinto-da-Rocha and Prendini, 2013 Chaerilobuthus Lourenço and Beigel, 2011 Chaerilus Simon, 1877 Chaneke Francke, Teruel and Santibáñez-López, 2014 Charmus Karsch, 1879 Cheloctonus Pocock, 1892 Chersonesometrus Couzijn, 1978 Chihuahuanus González-Santillán and Prendini, 2013 Chiromachetes Pocock, 1899 Chiromachus Pocock, 1893 Cicileiurus Teruel, 2007 Cicileus Vachon, 1948 Compsobuthus Vachon, 1949 Congobuthus Lourenço, 1999 Cretaceousbuthus Lourenço, 2022 Cretaceoushormiops Lourenço, 2018 Cretaceousopisthacanthus Lourenço in Lourenço and Velten, 2021 Cryptoiclus Teruel and Kovarík, 2012 Darchenia Vachon, 1977 Deccanometrus Prendini and Loria, 2020 Didymocentrus Kraepelin, 1905 Diplocentrus Peters, 1861 Egyptobuthus Lourenço, 1999 Electrochaerilus Santiago-Blay, Fet, Soleglad and Anderson, 2004 Euscorpius Thorell, 1876 Femtobuthus Lowe, 2010 Fetilinia Lowe and Kovarík, 2021 Franckeus Soleglad and Fet, 2005 Gertschius Graham and Soleglad, 2007 Gigantometrus Couzijn, 1978 Gint Kovarík, Lowe, Plísková and Stáhlavsky, 2013 Graemeloweus Soleglad, Fet, Graham and Ayrey, 2016 Grosphus Simon, 1880 Guyanochactas Lourenço, 1998 Hadogenes Kraepelin, 1894 Hadrurochactas Pocock, 1893 Hadruroides Pocock, 1893 Hadrurus Thorell, 1876 Hemibuthus Pocock, 1900 Hemilychas Hirst, 1911 Hemiscorpius Peters, 1861 Heteroctenus Pocock, 1893 Heterometrus Ehrenberg, 1828 Heteronebo Pocock, 1899 Heteroscorpion Birula, 1903 Himalayotityobuthus Lourenço, 1997 Hoffmannihadrurus Fet and Soleglad, 2004 Hormiops Fage, 1933 Hormurus Thorell, 1876 Hottentotta Birula, 1908 Iomachus Pocock, 1893 Iranobuthus Kovarík, 1997 Ischnotelson Esposito, Yamaguti, Souza, Pinto-da-Rocha and Prendini, 2017 Isometroides Keyserling, 1885 Isometrus Ehrenberg, 1828 Iurus Thorell, 1876 Jaguajir Esposito, Yamaguti, Souza, Pinto-da-Rocha and Prendini, 2017 Janalychas Kovarík, 2019 Javanimetrus Couzijn, 1981 Karasbergia Hewitt, 1913 Kochius Soleglad and Fet, 2008 Kolotl Santibañez-López, Francke and Prendini, 2014 Konetontli González-Santillán and Prendini, 2013 Kovarikia Soleglad, Fet and Graham, 2014 Kraepelinia Vachon, 1974 Kuarapu Francke and Ponce Saavedra, 2010 Langxie Tang, Jia and Liu, 2023 Lanzatus Kovarík, 2001 Leiurus Ehrenberg, 1828 Letoiurus Parmakelis et al., 2022 Liobuthus Birula, 1898 Liocheles Sundevall, 1833 Lisposoma Lawrence, 1928 Lissothus Vachon, 1948 Lychas C. L. Koch, 1845 Lychasioides Vachon, 1974 Maaykuyak González-Santillán and Prendini, 2013 Mauritanobuthus Qi and Lourenço, 2007 Mauryius Ojanguren-Affilastro and Mattoni, 2017 Megachactops Ochoa, Rojas-Runjaic, Pinto-da-Rocha and Prendini, 2013 Megacormus Karsch, 1881 Mesobuthus Vachon, 1950 Mesomexovis González-Santillán and Prendini, 2013 Mesotityus González-Sponga, 1981 Metaiurus Parmakelis et al., 2022 Microananteris Lourenço, 2003 Microbuthus Kraepelin, 1898 Microcharmus Lourenço, 1995 Microtityus Kjellesvig-Waering, 1966 Mioscorpio Kjellesvig-Waering, 1986 Nebo Simon, 1878 Neobuthus Hirst, 1911 Neocalchas Yagmur, Soleglad, Fet and Kovarík, 2013 Neochactas Soleglad and Fet, 2003 Neogrosphus Lourenço, 1995 Neoprotobuthus Lourenço, 2000 Nullibrotheas Williams, 1974 Odontobuthus Vachon, 1950 Odonturus Karsch, 1879 Oiclus Simon, 1880 Olivierus Farzanpay, 1987 Opisthacanthus Peters, 1861 Opistophthalmus C. L. Koch, 1837 Orobothriurus Maury, 1975 Orthochiroides Kovarík, 1998 Orthochirus Karsch, 1891 Pachakutej Ochoa, 2004 Palaeoakentrobuthus Lourenço and Weitschat, 2000 Palaeoananteris Lourenço and Weitschat, 2001 Palaeoburmesebuthus Lourenço, 2002 Palaeocheloctonus Lourenço, 1996 Palaeoeuscorpius Lourenço, 2003 Palaeogrosphus Lourenço, 2000 Palaeoisometrus Lourenço and Weitschat, 2005 Palaeolychas Lourenço and Weitschat, 1996 Palaeoprotobuthus Lourenço and Weitschat, 2000 Palaeospinobuthus Lourenço, Henderickx and Weitschat, 2005 Palaeotityobuthus Lourenço and Weitschat, 2000 Palaeotrilineatus Lourenço, 2012 Pandiborellius Rossi, 2015 Pandinoides Fet, 1997 Pandinops Birula, 1913 Pandinopsis Vachon, 1974 Pandinurus Fet, 1997 Pandinus Thorell, 1876 Pandipalpus Rossi, 2015 Pantobuthus Lourenço and Duhem, 2009 Parabuthus Pocock, 1890 Parascorpiops Banks, 1928 Paravaejovis Williams, 1980 Paruroctonus Werner, 1934 Pectinibuthus Fet in Orlov and Vasilyev, 1984 Phoniocercus Pocock, 1893 Physoctonus Mello-Leitão, 1934 Picobuthus Lowe, 2010 Plesiobuthus Pocock, 1900 Plesiochactas Pocock, 1900 Polisius Fet, Capes and Sissom, 2001 Protoiurus Soleglad, Fet, Kovarík and Yagmur, 2012 Pseudochactas Gromov, 1998 Pseudolissothus Lourenço, 2001 Pseudolychas Kraepelin, 1911 Pseudouroctonus Stahnke, 1974 Pseudouroplectes Lourenço, 1995 Qianxie Tang, 2022 Razianus Farzanpay, 1987 Reddyanus Vachon, 1972 Rhopalurus Thorell, 1876 Rugodentus Bastawade, Sureshan and Radhakrishnan, 2005 Rumikiru Ojanguren-Affilastro, Mattoni, Ochoa and Prendini, 2012 Saharobuthus Lourenço and Duhem, 2009 Sahyadrimetrus Prendini and Loria, 2020 Sardoscorpius Tropea and Onnis, 2020 Sassanidothus Farzanpay, 1987 Scorpio Linnaeus, 1758 Scorpiops Peters, 1861 Serradigitus Stahnke, 1974 Sinoscorpius Hong, 1983 Smeringurus Haradon, 1983 Somalibuthus Kovarík, 1998 Somalicharmus Kovarík, 1998 Sotanochactas Francke, 1986 Spelaeolychas Kovarík, 2019 Spinoburmesebuthus Lourenço in Lourenço and Velten, 2017 Spinochactas Lourenço, 2016 Srilankametrus Couzijn, 1981 Stahnkeus Soleglad and Fet, 2006 Stygochactas Vignoli and Prendini, 2009 Sucinlourencous Rossi, 2015 Superstitionia Stahnke, 1940 Syntropis Kraepelin, 1900 Tarsoporosus Francke, 1978 Tehuankea Cekalovic, 1973 Teruelius Lowe and Kovarík, 2019 Tetratrichobothrius Birula, 1917 Teuthraustes Simon, 1878 Thaicharmus Kovarík, 1995 Thestylus Simon, 1880 Thorellius Soleglad and Fet, 2008 Tibetiomachus Lourenço and Qi, 2006 Timogenes Simon, 1880 Tityobuthus Pocock, 1893 Tityopsis Armas, 1974 Tityus C. L. Koch, 1836 Troglocormus Francke, 1981 Troglokhammouanus Lourenço, 2007 Troglopolyphemos Rossi, 2018 Troglorhopalurus Lourenço, Baptista and Giupponi, 2004 Troglotayosicus Lourenço, 1981 Troglotityobuthus Lourenço, 2000 Trypanothacus Lowe, Kovarík, Stockmann and Stáhlavsky, 2019 Typhlochactas Mitchell, 1968 Uroctonites Williams and Savary, 1991 Uroctonus Thorell, 1876 Urodacus Peters, 1861 Urophonius Pocock, 1893 Uroplectes Peters, 1861 Vachonia Abalos, 1954 Vachoniochactas González-Sponga, 1978 Vachoniolus Levy, Amitai and Shulov, 1973 Vachonus Tikader and Bastawade, 1983 Vaejovis C. L. Koch, 1836 Vejovoidus Stahnke, 1974 Vietbocap Lourenço and Pham, 2010 Vizcaino González-Santillán and Prendini, 2013 Wernerius Soleglad and Fet, 2008 Xenobuthus Lowe, 2018 Zabius Thorell, 1893
Aegaeobuthus cyprius (Gantenbein and Kropf, 2000) Aegaeobuthus gallianoi (Ythier, 2018) Aegaeobuthus gibbosus (Brullé, 1832) Aegaeobuthus nigrocinctus (Ehrenberg, 1828) Aemngvantom lao (Lourenço, 2012) Aemngvantom thamnongpaseuam Prendini, Ehrenthal and Loria, 2021 Afroisometrus minshullae (Fitzpatrick, 1994) Afrolychas braueri (Kraepelin, 1896) Afrolychas burdoi (Simon, 1882) Akentrobuthus atakora Vignoli and Prendini, 2008 Akentrobuthus leleupi Lamoral, 1976 Akrav israchanani Levy, 2007 Alacran chamuco Francke, 2009 Alacran tartarus Francke, 1982 Alacran triquimera Santibañez-López, Francke and Prendini, 2014 Alayotityus delacruzi Armas, 1973 Alayotityus feti Teruel, 2004 Alayotityus granma Armas, 1984 Alayotityus juraguaensis Armas, 1973 Alayotityus lapidicola Teruel, 2002 Alayotityus nanus Armas, 1973 Alayotityus pallidus Teruel, 2002 Alayotityus sierramaestrae Armas, 1973 Alpiscorpius alpha (Caporiacco, 1950) Alpiscorpius beta (Caporiacco, 1950) Alpiscorpius caporiaccoi (Bonacina, 1980) Alpiscorpius croaticus (Caporiacco, 1950) Alpiscorpius delta Kovarík, Stundlová, Fet and Stáhlavsky, 2019 Alpiscorpius dinaricus (Caporiacco, 1950) Alpiscorpius gamma (Caporiacco, 1950) Alpiscorpius germanus (C. L. Koch, 1837) Alpiscorpius kappa Kovarík, Stundlová, Fet and Stáhlavsky, 2019 Alpiscorpius karamani Tropea, 2021 Alpiscorpius lambda Kovarík, Stundlová, Fet and Stáhlavsky, 2019 Alpiscorpius liburnicus Tvrtkovic and Rebrina in Podnar et al., 2022 Alpiscorpius mingrelicus (Kessler, 1874) Alpiscorpius omega Kovarík, Stundlová, Fet and Stáhlavsky, 2019 Alpiscorpius omikron Kovarík, Stundlová, Fet and Stáhlavsky, 2019 Alpiscorpius pavicevici Tropea, 2021 Alpiscorpius phrygius (Bonacina, 1980) Alpiscorpius sigma Kovarík, Stundlová, Fet and Stáhlavsky, 2019 Alpiscorpius uludagensis (Lacroix, 1995) Alpiscorpius ypsilon Kovarík, Stundlová, Fet and Stáhlavsky, 2019 Alpiscorpius zloporubovici Tropea, 2021 Ananteris arcadioi Botero-Trujillo, 2008 Ananteris ashaninka Kovarík, Teruel, Lowe and Friedrich, 2015 Ananteris ashmolei Lourenço, 1981 Ananteris asuncionensis González-Sponga, 2006 Ananteris balzanii Thorell, 1891 Ananteris barinensis González-Sponga, 2006 Ananteris bernabei Giupponi, Vasconcelos and Lourenço, 2009 Ananteris bianchinii Lourenço, Aguiar-Neto and Limeira-de-Oliveira, 2009 Ananteris bonito Lourenço, 2012 Ananteris cachimboensis Lourenço, Motta and da Silva, 2006 Ananteris camacan Lourenço, Giupponi and Leguin, 2013 Ananteris canalera Miranda and Armas, 2020 Ananteris capayaensis González-Sponga, 2006 Ananteris caracensis González-Sponga, 2006 Ananteris carrasco Lourenço and Motta, 2019 Ananteris catuaroi González-Sponga, 2006 Ananteris caucaguitensis González-Sponga, 2006 Ananteris chagasi Giupponi, Vasconcelos and Lourenço, 2009 Ananteris charlescorfieldi Lourenço, 2001 Ananteris chirimakei González-Sponga, 2006 Ananteris cisandinus Lourenço, 2015 Ananteris claviformis González-Sponga, 2006 Ananteris coineaui Lourenço, 1982 Ananteris columbiana Lourenço, 1991 Ananteris cryptozoicus Lourenço, 2005 Ananteris cumbensis González-Sponga, 2006 Ananteris curariensis González-Sponga, 2006 Ananteris cussinii Borelli, 1910 Ananteris dacostai Ythier, Chevalier and Lourenço, 2020 Ananteris dekeyseri Lourenço, 1982 Ananteris deniseae Lourenço, 1997 Ananteris desiderio Lourenço, Giupponi and Leguin, 2013 Ananteris diegorojasi Rojas-Runjaic, 2005 Ananteris dorae Botero-Trujillo, 2008 Ananteris ehrlichi Lourenço, 1994 Ananteris elguapoi González-Sponga, 2006 Ananteris elisabethae Lourenço, 2003 Ananteris evellynae Lourenço, 2004 Ananteris faguasi Botero-Trujillo, 2009 Ananteris festae Borelli, 1899 Ananteris franckei Lourenço, 1982 Ananteris gorgonae Lourenço and Flórez, 1989 Ananteris guiripaensis González-Sponga, 2006 Ananteris guyanensis Lourenço and Monod, 1999 Ananteris infuscata Lourenço, Giupponi and Leguin, 2013 Ananteris inoae González-Sponga, 2006 Ananteris intermedia Lourenço, 2012 Ananteris kalina Ythier, 2018 Ananteris karupina Lourenço, 2021 Ananteris kayapo Lourenço, 2022 Ananteris kuryi Giupponi, Vasconcelos and Lourenço, 2009 Ananteris leilae Lourenço, 1999 Ananteris luciae Lourenço, 1984 Ananteris madeirensis Lourenço and Duhem, 2010 Ananteris mamilihpan Ythier, Chevalier and Lourenço, 2020 Ananteris maniapurensis González-Sponga, 2006 Ananteris maranhensis Lourenço, 1987 Ananteris mariaelenae Lourenço, 1999 Ananteris mariaterezae Lourenço, 1982 Ananteris martensi Lourenço, 2021 Ananteris mauryi Lourenço, 1982 Ananteris meridana González-Sponga, 2006 Ananteris michaelae Lourenço, 2013 Ananteris myriamae Botero-Trujillo, 2007 Ananteris nairae Lourenço, 2004 Ananteris norae González-Sponga, 2006 Ananteris obscura Lourenço and Motta, 2021 Ananteris ochoai Botero-Trujillo and Flórez, 2011 Ananteris otavianoi Lira, Pordeus and Ribeiro de Albuquerque, 2017 Ananteris palmari Botero-Trujillo and Noriega, 2011 Ananteris paoensis González-Sponga, 2006 Ananteris paracotoensis González-Sponga, 2006 Ananteris pierrekondre Lourenço, Chevalier, Gangadin and Ythier, 2020 Ananteris plataensis González-Sponga, 2006 Ananteris platnicki Lourenço, 1993 Ananteris polleti Lourenço, 2016 Ananteris principalis González-Sponga, 2006 Ananteris pydanieli Lourenço, 1982 Ananteris riocaurensis González-Sponga, 2006 Ananteris riochicoi González-Sponga, 2006 Ananteris riomachensis Rojas-Runjaic, Portillo-Quintero and Borges, 2008 Ananteris roraima Lourenço and Duhem, 2010 Ananteris sabineae Lourenço, 2001 Ananteris sanchezi González-Sponga, 2006 Ananteris sepulvedai González-Sponga, 2006 Ananteris singularis González-Sponga, 2006 Ananteris sipilili Ythier, Chevalier and Lourenço, 2020 Ananteris solimariae Botero-Trujillo and Flórez, 2011 Ananteris surinamensis Lourenço, 2012 Ananteris terueli Kovarík, 2006 Ananteris tolimana Teruel and García, 2007 Ananteris tresor Ythier, Chevalier and Lourenço, 2020 Ananteris turumbanensis González-Sponga, 1980 Ananteris venezuelensis González-Sponga, 1972 Ananteris volschenski Botero-Trujillo, 2009 Ananteris zuliana González-Sponga, 2006 Ananteroides feae Borelli, 1911 Ananteroides inexpectatus Lourenço, 2013 Anatoliurus kraepelini (von Ubisch, 1922) Anatoliurus kumlutasi (Yagmur, Soleglad, Fet and Kovarík, 2015) Androctonus aeneas C. L. Koch, 1839 Androctonus afghanus Lourenço and Qi, 2006 Androctonus agrab Ythier and Lourenço, 2022 Androctonus aleksandrplotkini Lourenço and Qi, 2007 Androctonus amoreuxi (Audouin, 1826) Androctonus australis (Linnaeus, 1758) Androctonus baluchicus (Pocock, 1900) Androctonus barbouri (Werner, 1932) Androctonus bartolozzii Rossi and Merendino, 2016 Androctonus bicolor Ehrenberg, 1828 Androctonus bourdoni Vachon, 1948 Androctonus burkinensis Ythier, 2021 Androctonus cacahuati Lourenço, 2023 Androctonus cholistanus Kovarík and Ahmed, 2013 Androctonus crassicauda (Olivier, 1807) Androctonus dekeyseri Lourenço, 2005 Androctonus donairei Rossi, 2015 Androctonus eburneus (Pallary, 1928) Androctonus finitimus (Pocock, 1897) Androctonus gonneti Vachon, 1948 Androctonus hoggarensis (Pallary, 1929) Androctonus kunti Yagmur, 2023 Androctonus liouvillei (Pallary, 1924) Androctonus maelfaiti Lourenço, 2005 Androctonus maroccanus Lourenço, Ythier and Leguin, 2009 Androctonus mauritanicus (Pocock, 1902) Androctonus pallidus Lourenço, Duhem and Cloudsley-Thompson, 2012 Androctonus robustus Kovarík and Ahmed, 2013 Androctonus santi Lourenço, 2015 Androctonus sergenti Vachon, 1948 Androctonus simonettai Rossi, 2015 Androctonus sistanus Barahoei and Mirshamsi in Barahoei et al., 2022 Androctonus tenuissimus Teruel, Kovarík and Turiel, 2013 Androctonus tibesti Lourenço and El-Hennawy, 2022 Androctonus tigrai Lourenço, Rossi and Sadine, 2015 Androctonus tihamicus Alqahtani, Yagmur and Badry, 2023 Androctonus togolensis Lourenço, 2008 Androctonus tropeai Rossi, 2015 Androctonus turkiyensis Yagmur, 2021 Anomalobuthus krivochatskyi Teruel, Kovarík and Fet, 2018 Anomalobuthus lowei Teruel, Kovarík and Fet, 2018 Anomalobuthus pavlovskyi Teruel, Kovarík and Fet, 2018 Anomalobuthus rickmersi Kraepelin, 1900 Anomalobuthus talebii Teruel, Kovarík, Navidpour and Fet, 2014 Anomalobuthus zarudnyi (Birula, 1911) Antridiscalceatus trezzii (Vignoli and Kovarík, 2003) Anuroctonus phaiodactylus (Wood, 1863) Anuroctonus pococki Soleglad and Fet, 2004 Aops oncodactylus Volschenk and Prendini, 2008 Apistobuthus pterygocercus Finnegan, 1932 Apistobuthus susanae Lourenço, 1998 Archaeoananteroides maderai Lourenço in Lourenço and Velten, 2016 Archaeobuthus estephani Lourenço, 2001 Archaeoscorpiops cretacicus Lourenço, 2015 Australobuthus xerolimniorum Locket, 1990 Auyantepuia aluku Ythier, 2018 Auyantepuia amapaensis Lourenço and Qi, 2007 Auyantepuia aurum Ythier, 2018 Auyantepuia laurae Ythier, 2015 Auyantepuia royi Ythier, 2018 Auyantepuia surinamensis Lourenço and Duhem, 2010 Babycurus ansorgei Hirst, 1911 Babycurus brignolii Lourenço and Rossi, 2017 Babycurus buettneri Karsch, 1886 Babycurus centrurimorphus Karsch, 1886 Babycurus dunlopi Kovarík, Lowe, Seiter, Plísková and Stáhlavsky, 2015 Babycurus gigas Kraepelin, 1896 Babycurus jacksoni (Pocock, 1890) Babycurus kirki (Pocock, 1890) Babycurus melanicus Kovarík, 2000 Babycurus ochraceus Masi, 1912 Babycurus pictus Pocock, 1896 Babycurus solegladi Lourenço, 2005 Babycurus wituensis Kraepelin, 1913 Baloorthochirus becvari Kovarík, 1996 Balsateres cisnerosi (Ponce-Saavedra and Sissom, 2004) Barbaracurus borellii (Rossi, 2018) Barbaracurus exquisitus (Lowe, 2000) Barbaracurus feti Kovarík, Lowe, Stáhlavsky and Hurre, 2019 Barbaracurus kabateki Kovarík, Lowe, Stáhlavsky and Just, 2022 Barbaracurus prudenti (Lourenço, 2013) Barbaracurus sofomarensis (Kovarík, Lowe, Seiter, Plísková and Stáhlavsky, 2015) Barbaracurus somalicus (Hirst, 1907) Barbaracurus subpunctatus (Borelli, 1925) Barbaracurus ugartei (Kovarík, 2000) Barbaracurus winklerorum Kovarík, Lowe and Stáhlavsky, 2018 Barbaracurus zambonellii (Borelli, 1902) Belisarius ibericus Lourenço, 2015 Belisarius xambeui Simon, 1879 Betaburmesebuthus bellus Lourenço, 2016 Betaburmesebuthus bidentatus Lourenço, 2015 Betaburmesebuthus fleissneri Lourenço in Lourenço and Velten, 2016 Betaburmesebuthus fuscus Xuan, Cai and Huang, 2023 Betaburmesebuthus joergi Lourenço and Rossi, 2017 Betaburmesebuthus kobberti Lourenço in Lourenço and Beigel, 2015 Betaburmesebuthus larafleissnerae Lourenço in Lourenço and Velten, 2016 Betaburmesebuthus muelleri Lourenço, 2015 Betaburmesebuthus villosus Xuan, Cai and Huang, 2023 Bioculus caboensis (Stahnke, 1968) Bioculus cerralvensis Stahnke, 1968 Bioculus comondae Stahnke, 1968 Bioculus cruzensis Stahnke, 1968 Bioculus parvulus Martín-Frías, 2004 Birulatus astartiae Stathi and Lourenço, 2003 Birulatus haasi Vachon, 1974 Birulatus israelensis Lourenço, 2002 Birulatus jordanensis Lourenço et al., 2021 Bothriurus aguardente Santos-da-Silva, Carvalho and Brescovit, 2017 Bothriurus andorhinas Lourenço, 2023 Bothriurus araguayae Vellard, 1934 Bothriurus asper Pocock, 1893 Bothriurus bertae Abalos, 1955 Bothriurus bocki Kraepelin, 1911 Bothriurus bonariensis (C. L. Koch, 1842) Bothriurus buecherli San Martín, 1963 Bothriurus burmeisteri Kraepelin, 1894 Bothriurus ceii Ojanguren-Affilastro, 2007 Bothriurus cerradoensis Lourenço, Motta, Pitaluga de Godoi and Araujo, 2004 Bothriurus chacoensis Maury and Acosta, 1993 Bothriurus chilensis (Molina, 1782) Bothriurus cordubensis Acosta, 1995 Bothriurus coriaceus Pocock, 1893 Bothriurus delmari Santos-da-Silva, Carvalho and Brescovit, 2017 Bothriurus dumayi Cekalovic, 1974 Bothriurus flavidus Kraepelin, 1911 Bothriurus goiano Lovato, Anker and Lourenço, 2021 Bothriurus guarani Maury, 1984 Bothriurus huincul Mattoni, 2007 Bothriurus illudens Mello-Leitão, 1947 Bothriurus inermis Maury, 1981 Bothriurus jesuita Ojanguren-Affilastro, 2003 Bothriurus keyserlingi Pocock, 1893 Bothriurus maculatus Kraepelin, 1911 Bothriurus mistral Ojanguren-Affilastro, Mattoni, Alfaro and Pizarro-Araya, 2023 Bothriurus mochaensis Cekalovic, 1982 Bothriurus moojeni Mello-Leitão, 1945 Bothriurus nendai Ojanguren-Affilastro and García-Mauro, 2010 Bothriurus noa Maury, 1984 Bothriurus olaen Acosta, 1997 Bothriurus pampa Ojanguren-Affilastro, 2002 Bothriurus patagonicus Maury, 1968 Bothriurus pichicuy Mattoni, 2002 Bothriurus picunche Mattoni, 2002 Bothriurus pora Mattoni and Acosta, 2005 Bothriurus prospicuus Mello-Leitão, 1932 Bothriurus rochai Mello-Leitão, 1932 Bothriurus rochensis San Martín, 1965 Bothriurus rubescens Mello-Leitão, 1947 Bothriurus sanctaecrucis Mattoni, 2007 Bothriurus signatus Pocock, 1893 Bothriurus sooretamensis San Martín, 1966 Bothriurus trivittatus Werner, 1939 Bothriurus vachoni San Martín, 1968 Bothriurus vittatus (Guérin-Méneville, 1838) Bothriurus voyati Maury, 1973 Bothriurus wilhelmi Cekalovic, 1976 Bothriurus xingu Lourenço, 2016 Bothriurus ypsilon Mello-Leitão, 1935 Brachistosternus aconcagua Ojanguren-Affilastro and Scioscia, 2007 Brachistosternus alienus Lönnberg, 1898 Brachistosternus anandrovestigia Ojanguren-Affilastro, Pizarro-Araya and Ochoa, 2018 Brachistosternus andinus Chamberlin, 1916 Brachistosternus angustimanus Ojanguren Affilastro and Roig Alsina, 2001 Brachistosternus artigasi Cekalovic, 1974 Brachistosternus barrigai Ojanguren-Affilastro and Pizarro-Araya, 2014 Brachistosternus borellii Kraepelin, 1911 Brachistosternus castroi Mello-Leitão, 1940 Brachistosternus cekalovici Ojanguren-Affilastro, 2005 Brachistosternus cepedai Ojanguren-Affilastro, Agusto, Pizarro-Araya and Mattoni, 2007 Brachistosternus chango Ojanguren-Affilastro, Mattoni and Prendini, 2007 Brachistosternus chilensis Kraepelin, 1911 Brachistosternus chimba Ojanguren-Affilastro, Alfaro and Pizarro-Araya, 2021 Brachistosternus contisuyu Ojanguren-Affilastro, Pizarro-Araya and Ochoa, 2018 Brachistosternus coquimbo Ojanguren-Affilastro, Agusto, Pizarro-Araya and Mattoni, 2007 Brachistosternus diaguita Ojanguren-Affilastro in Ojanguren-Affilastro et al., 2023 Brachistosternus donosoi Cekalovic, 1974 Brachistosternus ehrenbergii (Gervais, 1841) Brachistosternus ferrugineus (Thorell, 1876) Brachistosternus galianoae Ojanguren-Affilastro, 2002 Brachistosternus gayi Ojanguren-Affilastro, Pizarro-Araya and Ochoa, 2018 Brachistosternus intermedius Lönnberg, 1902 Brachistosternus kamanchaca Ojanguren-Affilastro, Mattoni and Prendini, 2007 Brachistosternus kovariki Ojanguren-Affilastro, 2003 Brachistosternus llullaillaco Ojanguren-Affilastro, Alfaro and Pizarro-Araya, 2021 Brachistosternus mattonii Ojanguren-Affilastro, 2005 Brachistosternus misti Ojanguren-Affilastro, Pizarro-Araya and Ochoa, 2018 Brachistosternus montanus Roig Alsina, 1977 Brachistosternus multidentatus Maury, 1984 Brachistosternus negrei Cekalovic, 1975 Brachistosternus ninapo Ochoa, 2004 Brachistosternus ochoai Ojanguren-Affilastro, 2004 Brachistosternus paposo Ojanguren-Affilastro and Pizarro-Araya, 2014 Brachistosternus paulae Ojanguren-Affilastro, 2003 Brachistosternus pegnai Cekalovic, 1969 Brachistosternus pentheri Mello-Leitão, 1931 Brachistosternus perettii Ojanguren-Affilastro and Mattoni, 2006 Brachistosternus peruvianus Toledo Piza, 1974 Brachistosternus philippii Ojanguren-Affilastro, Pizarro-Araya and Ochoa, 2018 Brachistosternus piacentinii Ojanguren-Affilastro, 2003 Brachistosternus prendinii Ojanguren-Affilastro, 2003 Brachistosternus quiscapata Ochoa and Acosta, 2002 Brachistosternus roigalsinai Ojanguren Affilastro, 2002 Brachistosternus sciosciae Ojanguren Affilastro, 2002 Brachistosternus simonae Lourenço, 2000 Brachistosternus telteca Ojanguren Affilastro, 2000 Brachistosternus titicaca Ochoa and Acosta, 2002 Brachistosternus turpuq Ochoa, 2002 Brachistosternus weijenberghi (Thorell, 1877) Brachistosternus zambrunoi Ojanguren-Affilastro, 2002 Brazilobothriurus pantanalensis Lourenço and Monod, 2000 Broteochactas cauaburi Lourenço, Araujo and Franklin, 2010 Broteochactas cocuyensis González-Sponga, 2004 Broteochactas danielleae Lourenço, 2007 Broteochactas gollmeri (Karsch, 1879) Broteochactas manisapanensis (González-Sponga, 1992) Broteochactas mapuera Lourenço, 1988 Broteochactas mauriciodiasi Lourenço, 2017 Broteochactas niemeyerae Lourenço, Giupponi and Pedroso, 2011 Broteochactas nitidus Pocock, 1893 Broteochactas parimensis González-Sponga, 2004 Broteochactas porosus (Pocock, 1900) Broteochactas purus Lourenço, 2017 Broteochactas scorzai Dagert, 1957 Broteochactas silves Lourenço, 2014 Broteochactas venezuelensis (González-Sponga, 1996) Broteochactas verneti (González-Sponga, 1992) Broteochactas vestigialis (González-Sponga, 1978) Brotheas amazonicus Lourenço, 1988 Brotheas bolivianus Lourenço, 2008 Brotheas camposi González-Sponga, 1972 Brotheas caramaschii Lourenço, Giupponi and Pedroso, 2011 Brotheas cataniapensis González-Sponga, 1997 Brotheas cristinae Lourenço, 2007 Brotheas cunucunumensis González-Sponga, 1984 Brotheas dasilvai González-Sponga, 1978 Brotheas gervaisii Pocock, 1893 Brotheas granimanus Pocock, 1898 Brotheas granulatus Simon, 1877 Brotheas henriquesi Lourenço and Machado, 2004 Brotheas humboldti González-Sponga, 1980 Brotheas jourdani Lourenço, 1997 Brotheas libinallyi González-Sponga, 1978 Brotheas lichyi González-Sponga, 1980 Brotheas mawarinumensis González-Sponga, 1991 Brotheas mingueti González-Sponga, 1973 Brotheas munozi González-Sponga, 1997 Brotheas ocamoi González-Sponga, 2004 Brotheas overali Lourenço, 1988 Brotheas paraensis Simon, 1880 Brotheas perezramirezi González-Sponga, 1996 Brotheas rionegroensis González-Sponga, 1996 Brotheas sanabriai González-Sponga, 1997 Brotheas silvestris Lourenço, 1988 Brotheas tapajos Lourenço, 2012 Brotheas wareipai González-Sponga, 2004 Brotheas wilmeri González-Sponga, 1980 Burmesescorpiops groehni Lourenço, 2016 Buthacus agarwali Zambre and Lourenço, 2010 Buthacus ahaggar Lourenço, Kourim and Sadine, 2017 Buthacus algerianus Lourenço, 2006 Buthacus amitaii Cain, Gefen and Prendini, 2021 Buthacus arava Cain, Gefen and Prendini, 2021 Buthacus arenicola (Simon, 1885) Buthacus armasi Lourenço, 2013 Buthacus bicolor Afifeh, Al-Saraireh, Baker, Amr and Lourenço, 2022 Buthacus birulai Lourenço, 2006 Buthacus clevai Lourenço, 2001 Buthacus elmenia Lourenço and Sadine in Lourenço et al., 2017 Buthacus foleyi Vachon, 1948 Buthacus frontalis Werner, 1936 Buthacus fuscata Pallary, 1929 Buthacus golovatchi Lourenço, Duhem and Cloudsley-Thompson, 2012 Buthacus leptochelys (Ehrenberg, 1829) Buthacus levyi Cain, Gefen and Prendini, 2021 Buthacus macrocentrus (Ehrenberg, 1828) Buthacus mahraouii Lourenço, 2004 Buthacus maroccanus Lourenço, 2006 Buthacus nigerianus Lourenço and Qi, 2006 Buthacus nigroaculeatus Levy, Amitai and Shulov, 1973 Buthacus nitzani Levy, Amitai and Shulov, 1973 Buthacus occidentalis Vachon, 1953 Buthacus pakistanensis Lourenço and Qi, 2006 Buthacus sadinei Ythier, 2022 Buthacus samiae Lourenço and Sadine, 2015 Buthacus spatzi (Birula, 1911) Buthacus spinatus Lourenço, Bissati and Sadine, 2016 Buthacus stockmanni Kovarík, Lowe and Stáhlavsky, 2016 Buthacus striffleri Lourenço, 2004 Buthacus tadmorensis (Simon, 1892) Buthacus villiersi Vachon, 1949 Buthacus williamsi Lourenço and Leguin, 2009 Buthacus yotvatensis Levy, Amitai and Shulov, 1973 Buthacus ziegleri Lourenço, 2000 Butheoloides annieae Lourenço, 1986 Butheoloides aymerichi Lourenço, 2002 Butheoloides charlotteae Lourenço, 2000 Butheoloides cimrmani Kovarík, 2003 Butheoloides granulatus Lourenço, Duhem and Cloudsley-Thompson, 2012 Butheoloides grosseri Kovarík, 2016 Butheoloides hirsti Lourenço, 1996 Butheoloides littoralis Lourenço, Touloun and Boumezzough, 2011 Butheoloides maroccanus Hirst, 1925 Butheoloides milloti Vachon, 1948 Butheoloides monodi Vachon, 1950 Butheoloides nuer Kovarík, 2015 Butheoloides occidentalis Lourenço, Slimani and Berahou, 2003 Butheoloides polisi Lourenço, 1996 Butheoloides savanicola Lourenço, 2013 Butheoloides schwendingeri Lourenço, 2002 Butheoloides slimanii Lourenço, 2010 Butheoloides vanderberghi Lourenço, 2015 Butheoloides wilsoni Lourenço, 1995 Butheolus andersoni (Pocock, 1895) Butheolus gallagheri Vachon, 1980 Butheolus hallani Lourenço and Rossi, 2017 Butheolus harrisoni Lowe, 2018 Butheolus thalassinus Simon, 1882 Butheolus villosus Hendrixson, 2006 Buthiscus bicalcaratus Birula, 1905 Buthiscus ifoghas Ythier and Lourenço, 2023 Buthoscorpio chinnarensis Aswathi, Sureshan and Lourenço, 2015 Buthoscorpio indicus Lourenço, 2012 Buthoscorpio politus (Pocock, 1899) Buthoscorpio rayalensis Javed, Rao, Mirza, Sanap and Tampal, 2010 Buthoscorpio sarasinorum (Karsch, 1891) Buthus adrianae Rossi, 2013 Buthus ahaggar Ythier, Sadine, Haddadi and Lourenço, 2021 Buthus ajax (C. L. Koch, 1839) Buthus alacanti Teruel and Turiel, 2020 Buthus albengai Lourenço, 2003 Buthus amri Lourenço, Yagmur and Duhem, 2010 Buthus apiatus Lourenço, El Bouhissi and Sadine, 2020 Buthus atlantis Pocock, 1889 Buthus aures Lourenço and Sadine, 2016 Buthus awashensis Kovarík, 2011 Buthus baeticus Teruel and Turiel, 2020 Buthus balmensis Ythier and Laborieux, 2022 Buthus barcaeus Birula, 1909 Buthus berberensis Pocock, 1900 Buthus bobo Ythier, 2021 Buthus bonito Lourenço and Geniez, 2005 Buthus boumalenii Touloun and Boumezzough, 2011 Buthus boussaadi Lourenço, Chichi and Sadine, 2018 Buthus brignolii Lourenço, 2003 Buthus castellano Teruel and Turiel, 2022 Buthus centroafricanus Lourenço, 2016 Buthus chambiensis Kovarík, 2006 Buthus confluens Lourenço, Touloun and Boumezzough, 2012 Buthus danyii Rossi, 2017 Buthus delafuentei Teruel and Turiel, 2020 Buthus draa Lourenço and Slimani, 2004 Buthus dunlopi Kovarík, 2006 Buthus duprei Rossi and Tropea, 2016 Buthus egyptiensis Lourenço, 2012 Buthus elhennawyi Lourenço, 2005 Buthus elizabethae Lourenço, 2005 Buthus elmoutaouakili Lourenço and Qi, 2006 Buthus elongatus Rossi, 2012 Buthus gabani Ythier, 2021 Buthus garcialorcai Teruel and Turiel, 2020 Buthus goyffoni Abidi, Sadine and Lourenço in Abidi et al., 2021 Buthus halius (C. L. Koch, 1839) Buthus hassanini Lourenço, Duhem and Cloudsley-Thompson, 2012 Buthus iaspis Teruel and Turiel, 2022 Buthus ibericus Lourenço and Vachon, 2004 Buthus intermedius (Ehrenberg in Hemprich and Ehrenberg, 1829) Buthus intumescens (Ehrenberg in Hemprich and Ehrenberg, 1829) Buthus israelis Shulov and Amitai, 1959 Buthus jianxinae Lourenço, 2005 Buthus karoraensis Rossi and Tropea, 2016 Buthus kunti Yagmur, Koc and Lourenço, 2011 Buthus labuschagnei Lourenço, 2015 Buthus lienhardi Lourenço, 2003 Buthus lourencoi Rossi, Tropea and Yagmur, 2013 Buthus malhommei Vachon, 1949 Buthus manchego Teruel and Turiel, 2020 Buthus mardochei Simon, 1878 Buthus mariefranceae Lourenço, 2003 Buthus maroccanus Birula, 1903 Buthus montanus Lourenço and Vachon, 2004 Buthus nabataeus Lourenço, Afifeh and Al-Saraireh in Lourenço et al., 2021 Buthus nigrovesiculosus Hirst, 1925 Buthus occidentalis Lourenço, Sun and Zhu, 2009 Buthus occitanus (Amoreux, 1789) Buthus orientalis Lourenço and Simon, 2012 Buthus oudjanii Lourenço, 2017 Buthus paris (C. L. Koch, 1839) Buthus parroti Vachon, 1949 Buthus pedrosousai Teruel and Turiel, 2021 Buthus pococki Kovarík, Stáhlavsky and Elmi, 2020 Buthus prudenti Lourenço and Leguin, 2012 Buthus pusillus Lourenço, 2013 Buthus pyrenaeus Ythier, 2021 Buthus rochati Lourenço, 2003 Buthus saharicus Saddine, Bissati and Lourenço, 2016 Buthus serrano Teruel and Turiel, 2020 Buthus somalilandus Kovarík, Stáhlavsky and Elmi, 2020 Buthus tassili Lourenço, 2002 Buthus trinacrius Lourenço and Rossi, 2013 Buthus tunetanus (Herbst, 1800) Buthus turkana Ythier and Lourenço, 2022 Buthus yemenensis Lourenço, 2008 Buthus zeylensis Pocock, 1900 Calchas anlasi Yagmur, Soleglad, Fet and Kovarík, 2013 Calchas birulai Fet, Soleglad and Kovarík, 2009 Calchas kosswigi Yagmur, Soleglad, Fet and Kovarík, 2013 Calchas nordmanni Birula, 1899 Caraboctonus keyserlingi Pocock, 1893 Catalinia andreas (Gertsch and Soleglad, 1972) Catalinia ayreyi Teruel and Myers, 2019 Catalinia castanea (Gertsch and Soleglad, 1972) Catalinia minima (Kraepelin, 1911) Catalinia thompsoni (Gertsch and Soleglad, 1972) Cazierius alayoni Armas, 1999 Cazierius asper Teruel, 2006 Cazierius cayacoa Teruel, Carrero Jiménez and Santos, 2021 Cazierius chryseus Teruel and Armas, 2006 Cazierius ciguayo Teruel, Carrero Jiménez and Santos, 2021 Cazierius garridoi Armas, 2005 Cazierius granulosus Teruel, 2013 Cazierius gundlachii (Karsch, 1880) Cazierius neibae Kovarík and Teruel, 2014 Cazierius paradoxus Teruel and Diaz, 2004 Cazierius parvus Armas, 1984 Cazierius politus (Pocock, 1898) Cazierius torrei (Moreno, 1938) Centromachetes obscurus Mello-Leitão, 1932 Centromachetes pocockii (Kraepelin, 1894) Centromachetes titschaki (Werner, 1939) Centruroides alayoni Armas, 1999 Centruroides altagraciae Teruel, Armas and Kovarík, 2015 Centruroides anchorellus Armas, 1976 Centruroides arctimanus Armas, 1976 Centruroides baergi Hoffmann, 1932 Centruroides baldazoi Ponce-Saavedra, Linares-Guillén and Quijano-Ravell, 2022 Centruroides balsasensis Ponce-Saavedra and Francke, 2004 Centruroides bani Armas and Marcano Fondeur, 1987 Centruroides barbudensis (Pocock, 1898) Centruroides berstoni Goodman, Prendini, Francke and Esposito, 2021 Centruroides bertholdii (Thorell, 1876) Centruroides bicolor (Pocock, 1898) Centruroides bonito Quijano-Ravell, Teruel and Ponce-Saavedra, 2016 Centruroides caral Armas and Trujillo, 2013 Centruroides caribbeanus Teruel and Myers, 2017 Centruroides catemacoensis Goodman, Prendini, Francke and Esposito, 2021 Centruroides chamela Ponce-Saavedra and Francke, 2011 Centruroides chamulaensis Hoffmann, 1932 Centruroides chanae Goodman, Prendini, Francke and Esposito, 2021 Centruroides chiapanensis Hoffmann, 1932 Centruroides concordia Armas and Teruel, 2021 Centruroides cuauhmapan Goodman, Prendini, Francke and Esposito, 2021 Centruroides edwardsii (Gervais, 1843) Centruroides elegans (Thorell, 1876) Centruroides exilicauda (Wood, 1863) Centruroides exilimanus Teruel and Stockwell, 2002 Centruroides exsul (Meise, 1934) Centruroides fallassisimus Armas and Trujillo, 2010 Centruroides farri Armas, 1976 Centruroides flavopictus (Pocock, 1898) Centruroides franckei Santibañez-López and Contreras-Félix, 2013 Centruroides fulvipes (Pocock, 1898) Centruroides galano Teruel, 2001 Centruroides gracilis (Latreille, 1804) Centruroides granosus (Thorell, 1876) Centruroides griseus (C. L. Koch, 1844) Centruroides guanensis Franganillo, 1930 Centruroides hamadryas Goodman, Prendini, Francke and Esposito, 2021 Centruroides hentzi (Banks, 1900) Centruroides hirsuticauda Teruel, 2011 Centruroides hirsutipalpus Ponce-Saavedra and Francke, 2009 Centruroides hoffmanni Armas, 1996 Centruroides huichol Teruel, Ponce-Saavedra and Quijano-Ravell, 2015 Centruroides infamatus (C. L. Koch, 1844) Centruroides insulanus (Thorell, 1876) Centruroides ixil Trujillo and Armas, 2016 Centruroides jaragua Armas, 1999 Centruroides jorgeorum Santiago-Blay, 2009 Centruroides koesteri Kraepelin, 1912 Centruroides lauriadnae Ponce-Saavedra and Francke, 2019 Centruroides limbatus (Pocock, 1898) Centruroides limpidus (Karsch, 1879) Centruroides luceorum Armas, 1999 Centruroides lucidus Teruel, Armas and Kovarík, 2015 Centruroides marcanoi Armas, 1981 Centruroides margaritatus (Gervais, 1841) Centruroides mascota Ponce-Saavedra and Francke, 2011 Centruroides meisei Hoffmann, 1939 Centruroides melanodactylus Teruel, 2001 Centruroides navarroi Teruel, 2001 Centruroides nigrescens (Pocock, 1898) Centruroides nigrimanus (Pocock, 1898) Centruroides nigropunctatus Teruel, 2006 Centruroides nigrovariatus (Pocock, 1898) Centruroides nitidus (Thorell, 1876) Centruroides noxius Hoffmann, 1932 Centruroides ochraceus (Pocock, 1898) Centruroides orizaba Armas and Martín-Frías, 2003 Centruroides ornatus Pocock, 1902 Centruroides pallidiceps Pocock, 1902 Centruroides panamensis Quintero Arias and Esposito, 2014 Centruroides platnicki Armas, 1981 Centruroides pococki Sissom and Francke, 1983 Centruroides polito Teruel, 2007 Centruroides poncei Teruel, Kovarík, Baldazo-Monsivais and Hoferek, 2015 Centruroides possanii González-Santillán, Galán-Sánchez and Valdez-Velázquez, 2019 Centruroides rileyi Sissom, 1995 Centruroides robertoi Armas, 1976 Centruroides rodolfoi Santibañez-López and Contreras-Félix, 2013 Centruroides romeroi Quijano-Ravell, Armas, Francke and Ponce-Saavedra, 2019 Centruroides ruana Quijano-Ravell and Ponce-Saavedra, 2016 Centruroides sanandres Armas, Luna and Flórez, 2012 Centruroides sasae Santiago-Blay, 2009 Centruroides schmidti Sissom, 1995 Centruroides sculpturatus Ewing, 1928 Centruroides serrano Santibañez-López and Ponce-Saavedra, 2009 Centruroides simplex (Thorell, 1876) Centruroides sissomi Armas, 1996 Centruroides spectatus Teruel, 2006 Centruroides stockwelli Teruel, 2001 Centruroides suffusus Pocock, 1902 Centruroides tapachulaensis Hoffmann, 1932 Centruroides tecomanus Hoffmann, 1932 Centruroides tenuis (Thorell, 1876) Centruroides testaceus (De Geer, 1778) Centruroides thorellii (Kraepelin, 1891) Centruroides tuxtla Armas, 1999 Centruroides underwoodi Armas, 1976 Centruroides villegasi Baldazo-Monsivaiz, Ponce-Saavedra and Flores-Moreno, 2013 Centruroides vittatus (Say, 1821) Centruroides yucatanensis Goodman, Prendini, Francke and Esposito, 2021 Cercophonius granulosus Kraepelin, 1908 Cercophonius kershawi Glauert, 1930 Cercophonius michaelseni Kraepelin, 1908 Cercophonius queenslandae Acosta, 1990 Cercophonius squama (Gervais, 1843) Cercophonius sulcatus Kraepelin, 1908 Chactas adornellae Rossi, 2014 Chactas aequinoctialis (Karsch, 1879) Chactas alarconi González-Sponga, 2003 Chactas barbacoensis González-Sponga, 1987 Chactas barravierai Lourenço, 1997 Chactas braziliensis Lourenço, Aguiar and Franklin, 2005 Chactas brevicaudatus (Karsch, 1879) Chactas brownelli Lourenço, 1997 Chactas campoeliasensis González-Sponga, 2006 Chactas chabasquensis González-Sponga and Wall-González, 2007 Chactas choroniensis González-Sponga, 1978 Chactas chrysopus Pocock, 1893 Chactas exsul (Werner, 1939) Chactas ferruginosus González-Sponga, 1984 Chactas gansi González-Sponga, 1974 Chactas gestroi Kraepelin, 1912 Chactas granulosus González-Sponga, 2007 Chactas guinandcortesi González-Sponga, 2003 Chactas hatilloensis González-Sponga, 2007 Chactas hauseri Lourenço, 1997 Chactas interpuncta González-Sponga, 1987 Chactas iutensis González-Sponga, 2008 Chactas karschii Pocock, 1893 Chactas keyserlingii Pocock, 1893 Chactas koepckei Lourenço and Dastych, 2001 Chactas laevipes (Karsch, 1879) Chactas latuffi González-Sponga, 1976 Chactas lepturus Thorell, 1876 Chactas mahnerti Lourenço, 1995 Chactas maimirensis González-Sponga and Wall-González, 2007 Chactas major Kraepelin, 1912 Chactas marinae González-Sponga, 1987 Chactas mauriesi Lourenço and Flórez, 1990 Chactas moreti Lourenço, 2014 Chactas oxfordi González-Sponga, 1978 Chactas ozendai Lourenço, 1999 Chactas platillonensis González-Sponga and Wall-González, 2007 Chactas pleistocenicus Lourenço and Weitschat, 2005 Chactas raymondhansi Francke and Boos, 1986 Chactas reticulatus Kraepelin, 1912 Chactas rogelioi González-Sponga, 1972 Chactas rubrolineatus Simon, 1880 Chactas scabrimanus Kraepelin, 1912 Chactas setosus Kraepelin, 1912 Chactas simonii Pocock, 1893 Chactas tumaquensis González-Sponga and Wall-González, 2007 Chactas turguaensis González-Sponga, 2007 Chactas vanbenedenii (Gervais, 1843) Chactas venegasi González-Sponga, 2008 Chactas viloriai Rojas-Runjaic, 2004 Chactas yaupi Lourenço, 2014 Chactas yupai González-Sponga, 1994 Chactopsis amazonica Lourenço and Francke, 1986 Chactopsis barajuri González-Sponga, 1982 Chactopsis buhrnheimi Lourenço, 2003 Chactopsis chullachaqui Ochoa, Rojas-Runjaic, Pinto-da-Rocha and Prendini, 2013 Chactopsis curupira Ochoa, Rojas-Runjaic, Pinto-da-Rocha and Prendini, 2013 Chactopsis insignis Kraepelin, 1912 Chactopsis siapaensis González-Sponga, 1991 Chactopsis sujirima González-Sponga, 1982 Chactopsoides anduzei (González-Sponga, 1982) Chactopsoides gonzalezspongai Ochoa, Rojas-Runjaic, Pinto-da-Rocha and Prendini, 2013 Chactopsoides marahuacaensis (González-Sponga, 2004) Chactopsoides yanomami (Lourenço, Giupponi and Pedroso, 2011) Chaerilobuthus birmanicus Lourenço, 2015 Chaerilobuthus brandti Lourenço in Lourenço and Velten, 2022 Chaerilobuthus bruckschi Lourenço, 2015 Chaerilobuthus complexus Lourenço and Beigel, 2011 Chaerilobuthus enigmaticus Lourenço, 2015 Chaerilobuthus gigantosternum Lourenço, 2016 Chaerilobuthus knodelorum Lourenço, 2018 Chaerilobuthus meggeri Lourenço in Lourenço and Velten, 2021 Chaerilobuthus schwarzi Lourenço in Lourenço and Velten, 2015 Chaerilobuthus serratus Lourenço, 2016 Chaerilus agilis Pocock, 1899 Chaerilus agnellivanniorum Lourenço and Rossi, 2019 Chaerilus alberti Kovarík, 2019 Chaerilus andamanensis Lourenço, Duhem and Leguin, 2011 Chaerilus annapurna Lourenço and Duhem, 2010 Chaerilus assamensis Kraepelin, 1913 Chaerilus cavernicola Pocock, 1894 Chaerilus celebensis Pocock, 1894 Chaerilus ceylonensis Pocock, 1894 Chaerilus chapmani Vachon and Lourenço, 1985 Chaerilus chubluk Lourenço, Tran and Pham, 2020 Chaerilus cimrmani Kovarík, 2012 Chaerilus conchiformus Zhu, Han and Lourenço, 2008 Chaerilus granulatus Kovarík, Lowe, Hoferek, Forman and Král, 2015 Chaerilus hofereki Kovarík, Král, Korinková and Reyes Lerma, 2014 Chaerilus honba Lourenço, 2019 Chaerilus insignis Pocock, 1894 Chaerilus julietteae Lourenço, 2011 Chaerilus kampuchea Lourenço, 2012 Chaerilus kautti Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Chaerilus laevimanus Pocock, 1899 Chaerilus laoticus Lourenço and Zhu, 2008 Chaerilus lehtrarensis Khatoon, 1999 Chaerilus longimanus Kovarík and Lowe in Kovarík et al., 2015 Chaerilus mainlingensis Lourenço and Zhu, 2008 Chaerilus majkusi Kovarík, Lowe and Stáhlavsky, 2018 Chaerilus merendinoi Rossi, 2018 Chaerilus neradorum Kovarík, Lowe and Stáhlavsky, 2018 Chaerilus ojangureni Kovarík, 2005 Chaerilus pathom Lourenço and Pham, 2014 Chaerilus petrzelkai Kovarík, 2000 Chaerilus phami Lourenço, 2011 Chaerilus philippinus Lourenço and Ythier, 2008 Chaerilus pictus (Pocock, 1890) Chaerilus pseudoconchiformus Yin, Qiu, Pan, Li and Di, 2015 Chaerilus pulcherrimus Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Chaerilus rectimanus Pocock, 1899 Chaerilus robinsoni Hirst, 1911 Chaerilus sabinae Lourenço, 1995 Chaerilus seiteri Kovarík, 2012 Chaerilus sejnai Kovarík, 2005 Chaerilus solegladi Kovarík, 2012 Chaerilus spinatus Lourenço and Duhem, 2010 Chaerilus stockmannorum Kovarík, Lowe and Stáhlavsky, 2018 Chaerilus telnovi Lourenço, 2009 Chaerilus tessellatus Qi, Zhu and Lourenço, 2005 Chaerilus thai Lourenço, Sun and Zhu, 2010 Chaerilus tichyi Kovarík, 2000 Chaerilus tricostatus Pocock, 1899 Chaerilus truncatus Karsch, 1879 Chaerilus tryznai Kovarík, 2000 Chaerilus variegatus Simon, 1877 Chaerilus vietnamicus Lourenço and Zhu, 2008 Chaerilus wrzecionkoi Kovarík, 2012 Chaneke aliciae (Armas and Martín-Frías, 1998) Chaneke baldazoi Kovarík, Teruel and Lowe, 2016 Chaneke fogoso Francke, Teruel and Santibáñez-López, 2014 Chaneke hofereki Kovarík, Teruel and Lowe, 2016 Charmus brignolii Lourenço, 2000 Charmus indicus Hirst, 1915 Charmus laneus Karsch, 1879 Charmus saradieli Kovarík, Lowe, Ranawana, Hoferek and Jayarathne in Kovarík et al., 2016 Charmus sinhagadensis Tikader and Bastawade, 1983 Cheloctonus anthracinus Pocock, 1899 Cheloctonus crassimanus (Pocock, 1896) Cheloctonus glaber Kraepelin, 1896 Cheloctonus intermedius Hewitt, 1912 Cheloctonus jonesii Pocock, 1892 Cheloctonus kakongo Lourenço, 2022 Chersonesometrus bastawadei Prendini and Loria, 2020 Chersonesometrus beccaloniae (Kovarík, 2004) Chersonesometrus fulvipes (C. L. Koch, 1837) Chersonesometrus hendersoni Prendini and Loria, 2020 Chersonesometrus madraspatensis (Pocock, 1900) Chersonesometrus nathanorum Prendini and Loria, 2020 Chersonesometrus pelekomanus (Couzijn, 1981) Chersonesometrus shivashankari Prendini and Loria, 2020 Chersonesometrus tristis (Henderson, 1919) Chersonesometrus wroughtoni (Pocock, 1899) Chihuahuanus bilineatus (Pocock, 1898) Chihuahuanus cazieri (Williams, 1968) Chihuahuanus coahuilae (Williams, 1968) Chihuahuanus crassimanus (Pocock, 1898) Chihuahuanus glabrimanus (Sissom and Hendrixson, 2005) Chihuahuanus globosus (Borelli, 1915) Chihuahuanus kovariki (Soleglad and Fet, 2008) Chihuahuanus russelli (Williams, 1971) Chiromachetes agasthyamalaiensis Khandekar, Thackeray, Pawar, Gangalmale and Waghe, 2022 Chiromachetes fergusoni Pocock, 1899 Chiromachetes parakrami Sulakhe, Deshpande, Dandekar, Ketkar, Gowande, Padhye and Bastawade, 2020 Chiromachetes ramdasswamii Sulakhe, Deshpande, Dandekar, Ketkar, Gowande, Padhye and Bastawade, 2020 Chiromachetes sahyadriensis Mirza, Sanap and Zambre, 2015 Chiromachetes tirupati Lourenço, 1997 Chiromachus ochropus (C. L. Koch, 1837) Cicileiurus monticola Teruel, 2007 Cicileus cloudsleythompsoni Lourenço, 1999 Cicileus exilis (Pallary, 1928) Cicileus hoggarensis Lourenço and Rossi, 2015 Cicileus latellai Lourenço and Rossi, 2015 Cicileus montanus Lourenço and Rossi, 2015 Compsobuthus abyssinicus (Birula, 1903) Compsobuthus acutecarinatus (Simon, 1882) Compsobuthus afghanus Kovarík and Ahmed, 2007 Compsobuthus air Lourenço and Rossi, 2018 Compsobuthus andresi Lourenço, 2004 Compsobuthus arabicus Levy, Amitai and Shulov, 1973 Compsobuthus armenicus Lourenço and Leguin, 2015 Compsobuthus atrostriatus (Pocock, 1897) Compsobuthus berlandi Vachon, 1950 Compsobuthus birulai Lourenço, Leguin and Duhem, 2010 Compsobuthus boucheti Lourenço, Duhem and Cloudsley-Thompson, 2012 Compsobuthus brevimanus (Werner, 1936) Compsobuthus carmelitis Levy, Amitai and Shulov, 1973 Compsobuthus egyptiensis Lourenço, Sun and Zhu, 2009 Compsobuthus eritreaensis Kovarík, Lowe, Plísková and Stáhlavsky, 2016 Compsobuthus fuscatus Hendrixson, 2006 Compsobuthus garyi Lourenço and Vachon, 2001 Compsobuthus humaae Amir, Kamaluddin and Khan, 2005 Compsobuthus jakesi Kovarík, 2003 Compsobuthus jordanensis Levy, Amitai and Shulov, 1973 Compsobuthus kabateki Kovarík, 2003 Compsobuthus kafkai Kovarík, 2003 Compsobuthus kaftani Kovarík, 2003 Compsobuthus khaybari Afifeh, Aloufi and Al-Saraireh in Afifeh et al., 2021 Compsobuthus klaptoczi (Birula, 1909) Compsobuthus krali Kovarík, 2012 Compsobuthus levyi Kovarík, 2012 Compsobuthus longipalpis Levy, Amitai and Shulov, 1973 Compsobuthus mahazat Ythier and Lourenço, 2023 Compsobuthus maidensis Kovarík, 2018 Compsobuthus maindroni (Kraepelin, 1901) Compsobuthus manzonii (Borelli, 1915) Compsobuthus matthiesseni (Birula, 1905) Compsobuthus nematodactylus Lowe, 2009 Compsobuthus pakistanus Kovarík and Ahmed, 2007 Compsobuthus pallidus Hendrixson, 2006 Compsobuthus persicus Navidpour, Soleglad, Fet and Kovarík, 2008 Compsobuthus petriolii Vignoli, 2005 Compsobuthus plutenkoi Kovarík, 2003 Compsobuthus polisi Lowe, 2001 Compsobuthus rugosulus (Pocock, 1900) Compsobuthus satpuraensis Waghe, Gangalmale and Khandekar, 2022 Compsobuthus schmiedeknechti Vachon, 1949 Compsobuthus seicherti Kovarík, 2003 Compsobuthus setosus Hendrixson, 2006 Compsobuthus simoni Lourenço, 1999 Compsobuthus sindicus Kovarík and Ahmed, 2011 Compsobuthus sobotniki Kovarík, 2003 Compsobuthus somalilandus Kovarík, 2012 Compsobuthus tassili Lourenço, 2010 Compsobuthus tofti Lourenço, 2001 Compsobuthus tombouctou Lourenço, 2009 Compsobuthus turieli Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Compsobuthus ullrichi Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Compsobuthus vachoni Sissom, 1994 Compsobuthus vannii Rossi, 2017 Compsobuthus werneri (Birula, 1908) Congobuthus fagei Lourenço, 1999 Cretaceousbuthus fraaijeorum Lourenço in Lourenço and Velten, 2022 Cretaceousbuthus petersi Lourenço in Lourenço and Velten, 2023 Cretaceoushormiops knodeli Lourenço, 2018 Cretaceoushormiops staxi Lourenço in Lourenço and Velten, 2022 Cretaceousopisthacanthus smeelei Lourenço in Lourenço and Velten, 2021 Cryptoiclus rodriguezi Teruel and Kovarík, 2012 Darchenia bernadettae Vachon, 1977 Deccanometrus bengalensis (C. L. Koch, 1841) Deccanometrus latimanus (Pocock, 1894) Deccanometrus liurus (Pocock, 1897) Deccanometrus obscurus (Couzijn, 1981) Deccanometrus phipsoni (Pocock, 1893) Deccanometrus ubicki (Kovarík, 2004) Deccanometrus xanthopus (Pocock, 1897) Didymocentrus armasi Teruel and Rodríguez, 2008 Didymocentrus hasethi (Kraepelin, 1896) Didymocentrus hummelincki Francke, 1978 Didymocentrus jaumei Armas, 1976 Didymocentrus krausi Francke, 1978 Didymocentrus lesueurii (Gervais, 1844) Didymocentrus martinicae Teruel and Questel, 2020 Didymocentrus minor Francke, 1978 Didymocentrus nitidus (Hirst, 1907) Didymocentrus sanfelipensis Armas, 1976 Didymocentrus trinitarius (Franganillo, 1930) Didymocentrus vincenzoi Rossi, 2018 Didymocentrus waeringi Francke, 1978 Diplocentrus actun Armas and Palacios-Vargas, 2002 Diplocentrus bellator Teruel, 2003 Diplocentrus bereai Armas and Martín-Frías, 2004 Diplocentrus bicolor Contreras-Félix and Santibáñez-López, 2011 Diplocentrus chiapasensis Beutelspacher and Armas, 1998 Diplocentrus chol Francke, 2007 Diplocentrus churumuco Francke and Ponce Saavedra, 2005 Diplocentrus coddingtoni Stockwell, 1988 Diplocentrus colwelli Sissom, 1986 Diplocentrus coylei Fritts and Sissom, 1996 Diplocentrus cozumel Beutelspacher and Armas, 1998 Diplocentrus cueva Francke, 1978 Diplocentrus diablo Stockwell and Nilsson, 1987 Diplocentrus duende Santibañez-López and González-Santillán, 2017 Diplocentrus ferrugineus Fritts and Sissom, 1996 Diplocentrus formosus Armas and Martín-Frías, 2003 Diplocentrus franckei Santibáñez-López, 2014 Diplocentrus gertschi Sissom and Walker, 1992 Diplocentrus gladiator Beutelspacher and Trujillo, 1999 Diplocentrus hoffmanni Francke, 1977 Diplocentrus insularis Sagastume-Espinoza, Longhorn and Santibáñez-López, 2015 Diplocentrus izabal Armas and Trujillo, 2016 Diplocentrus jaca Armas and Martín-Frías, 2000 Diplocentrus keyserlingii Karsch, 1880 Diplocentrus kraepelini Santibañez-López, Francke and Prendini, 2013 Diplocentrus lachua Armas, Trujillo and Agreda, 2011 Diplocentrus landelinoi Trujillo and Armas, 2012 Diplocentrus leptomanus Villa-Corella, Silva-Kurumiya, Barrales-Alcalá, Van Devender and Francke, 2023 Diplocentrus lindo Stockwell and Baldwin, 2001 Diplocentrus longimanus Santibañez-López, Francke and Athanasiadis, 2011 Diplocentrus lourencoi Stockwell, 1988 Diplocentrus lucidus Stockwell, 1988 Diplocentrus luisae Guijosa, 1973 Diplocentrus majahuensis Baldazo-Monsivaiz, 2003 Diplocentrus maya Francke, 1977 Diplocentrus melici Armas, Martín-Frías and Berea, 2004 Diplocentrus mexicanus Peters, 1861 Diplocentrus mitchelli Francke, 1977 Diplocentrus mitlae Francke, 1977 Diplocentrus montecristo Armas and Martín-Frías, 2000 Diplocentrus motagua Armas and Trujillo, 2009 Diplocentrus ochoterenai Hoffmann, 1931 Diplocentrus ornatus Stockwell, 1988 Diplocentrus oxlajujbaktun Trujillo and Armas, 2012 Diplocentrus peloncillensis Francke, 1975 Diplocentrus perezi Sissom, 1991 Diplocentrus popti Trujillo and Armas, 2016 Diplocentrus rectimanus Pocock, 1898 Diplocentrus reddelli Francke, 1977 Diplocentrus roo Armas and Martín-Frías, 2005 Diplocentrus sagittipalpus Santibañez-López, Francke and Prendini, 2013 Diplocentrus santiagoi Stockwell, 1988 Diplocentrus silanesi Armas and Martín-Frías, 2000 Diplocentrus sinaan Armas and Martín-Frías, 2000 Diplocentrus sissomi Santibañez-López, Francke and Prendini, 2013 Diplocentrus spitzeri Stahnke, 1970 Diplocentrus steeleae Stockwell, 1988 Diplocentrus taibeli (Caporiacco, 1938) Diplocentrus tehuacanus Hoffmann, 1931 Diplocentrus tehuano Francke, 1977 Diplocentrus tenango Santibañez-López and Francke, 2008 Diplocentrus whitei (Gervais, 1844) Diplocentrus williamsi Sissom and Wheeler, 1995 Diplocentrus zacatecanus Hoffmann, 1931 Egyptobuthus vaissadei Lourenço, 1999 Electrochaerilus buckleyi Santiago-Blay, Fet, Soleglad and Anderson, 2004 Euscorpius aladaglarensis Tropea and Yagmur, 2016 Euscorpius alanyaensis Tropea, Yagmur, Parmakelis and Kunt, 2016 Euscorpius altadonnai Tropea, 2017 Euscorpius amorgensis Tropea, Fet, Parmakelis, Kotsakiozi and Stathi, 2017 Euscorpius aquilejensis (C. L. Koch, 1837) Euscorpius arikani Yagmur and Tropea, 2015 Euscorpius avcii Tropea, Yagmur, Koc, Yesilyurt and Rossi, 2012 Euscorpius balearicus Caporiacco, 1950 Euscorpius biokovensis Tropea and Ozimec, 2020 Euscorpius birulai Fet, Soleglad, Parmakelis, Kotsakiozi and Stathi, 2014 Euscorpius bonacinai Kovarík and Stáhlavsky, 2020 Euscorpius borovaglavaensis Tropea, 2015 Euscorpius calabriae Caporiacco, 1950 Euscorpius candiota Birula, 1903 Euscorpius canestrinii (Fanzago, 1872) Euscorpius carpathicus (Linnaeus, 1767) Euscorpius celanus Tropea, 2012 Euscorpius ciliciensis Birula, 1898 Euscorpius concinnus (C. L. Koch, 1837) Euscorpius corcyraeus Tropea and Rossi, 2012 Euscorpius curcici Tropea, Fet, Parmakelis, Kotsakiozi and Stathi, 2017 Euscorpius deltshevi Fet, Graham, Webber and Blagoev, 2014 Euscorpius drenskii Tropea, Fet, Parmakelis, Kotsakiozi and Stathi, 2015 Euscorpius erymanthius Tropea, Fet, Parmakelis, Kotsakiozi and Stathi, 2013 Euscorpius eskisehirensis Tropea and Yagmur, 2015 Euscorpius feti Tropea, 2013 Euscorpius garganicus Di Caporiacco, 1950 Euscorpius giachinoi Tropea and Fet, 2015 Euscorpius gocmeni Tropea, Yagmur and Yesilyurt, 2014 Euscorpius hadzii Caporiacco, 1950 Euscorpius hakani Tropea and Yagmur, 2016 Euscorpius honazicus Tropea, Yagmur, Karampatsou, Parmakelis and Yesilyurt, 2016 Euscorpius hyblaeus Tropea, 2016 Euscorpius idaeus Yagmur and Tropea, 2017 Euscorpius italicus (Herbst, 1800) Euscorpius janstai Kovarík and Stáhlavsky, 2020 Euscorpius kabateki Kovarík and Stáhlavsky, 2020 Euscorpius kinzelbachi Tropea, Fet, Parmakelis, Kotsakiozi and Stathi, 2014 Euscorpius koci Tropea and Yagmur, 2015 Euscorpius koschewnikowi Birula, 1900 Euscorpius kritscheri Fet, Soleglad, Parmakelis, Kotsakiozi and Stathi, 2013 Euscorpius lagostae Caporiacco, 1950 Euscorpius latinus Tropea and Parmakelis, 2022 Euscorpius lesbiacus Tropea et al., 2020 Euscorpius lycius Yagmur, Tropea and Yesilyurt, 2013 Euscorpius mylonasi Fet, Soleglad, Parmakelis, Kotsakiozi and Stathi, 2014 Euscorpius naupliensis (C. L. Koch, 1837) Euscorpius niciensis (C. L. Koch, 1841) Euscorpius oglasae Caporiacco, 1950 Euscorpius ossae Caporiacco, 1950 Euscorpius parthenopeius Tropea, Parmakelis, Sziszkosz, Balanika and Bouderka, 2014 Euscorpius popovi Tropea, Fet, Parmakelis, Kotsakiozi and Stathi, 2015 Euscorpius sadileki Kovarík and Stáhlavsky, 2020 Euscorpius salentinus Tropea, 2017 Euscorpius scaber Birula, 1900 Euscorpius scheraboni Kovarík and Stáhlavsky, 2020 Euscorpius sicanus (C. L. Koch, 1837) Euscorpius simaiakisi Tropea, Fet, Parmakelis and Stathi, 2022 Euscorpius solegladi Fet, Graham, Webber and Blagoev, 2014 Euscorpius stahlavskyi Tropea, Fet, Parmakelis, Kotsakiozi and Stathi, 2014 Euscorpius stefaniae Tropea and Parmakelis, 2022 Euscorpius studentium Karaman, 2020 Euscorpius sultanensis Tropea and Yagmur, 2016 Euscorpius tauricus (C. L. Koch, 1837) Euscorpius tergestinus (C. L. Koch, 1837) Euscorpius thracicus Kovarík, Lowe, Byronová and Stáhlavsky, 2020 Euscorpius trejaensis Tropea and Parmakelis, 2022 Euscorpius triantisi Tropea, Fet, Parmakelis and Stathi, 2022 Euscorpius vailatii Tropea and Fet, 2015 Euscorpius vignai Tropea, Fet, Parmakelis, Kotsakiozi and Stathi, 2014 Euscorpius yagmuri Kovarík, Fet and Soleglad, 2014 Femtobuthus shutuae Lowe, 2010 Fetilinia dentator Lowe and Kovarík, 2021 Franckeus kochi (Sissom, 1991) Franckeus minckleyi (Williams, 1968) Franckeus nitidulus (C. L. Koch, 1843) Franckeus peninsularis (Williams, 1980) Franckeus platnicki (Sissom, 1991) Franckeus rubrimanus (Sissom, 1991) Gertschius agilis (Sissom and Stockwell, 1991) Gertschius crassicorpus Graham and Soleglad, 2007 Gigantometrus swammerdami (Simon, 1872) Gigantometrus titanicus (Couzijn, 1981) Gint amoudensis Kovarík, Lowe, Just, Awale, Elmi and Stáhlavsky, 2018 Gint banfasae Kovarík and Lowe, 2019 Gint calviceps (Pocock, 1900) Gint childsi Kovarík, 2018 Gint dabakalo Kovarík and Mazuch, 2015 Gint gaitako Kovarík, Lowe, Plísková and Stáhlavsky, 2013 Gint gubanensis Kovarík, Lowe, Just, Awale, Elmi and Stáhlavsky, 2018 Gint insolitus (Borelli, 1925) Gint maidensis Kovarík, Lowe, Just, Awale, Elmi and Stáhlavsky, 2018 Gint marialuisae Rossi, 2015 Gint monicae Rossi, 2015 Gint puntlandus Kovarík and Mazuch, 2015 Graemeloweus glimmei (Hjelle, 1972) Graemeloweus iviei (Gertsch and Soleglad, 1972) Graemeloweus maidu (Savary and Bryson, 2016) Grosphus ambre Lourenço, Wilmé and Waeber, 2018 Grosphus angulatus Lowe and Kovarík, 2022 Grosphus darainensis Lourenço, Goodman and Ramilijaona, 2004 Grosphus goudoti Lourenço and Goodman, 2006 Grosphus halleuxi Lourenço, Wilmé, Soarimalala and Waeber, 2017 Grosphus hirtus Kraepelin, 1900 Grosphus madagascariensis (Gervais, 1843) Grosphus mandena Lourenço, 2005 Grosphus mayottensis Lourenço and Goodman, 2009 Grosphus polskyi Lourenço, Qi and Goodman, 2007 Grosphus rakotoariveloi Lourenço, Wilmé, Soarimalala and Waeber, 2017 Grosphus simoni Lourenço, Goodman and Ramilijaona, 2004 Grosphus tavaratra Lourenço, Soarimalala and Goodman, 2009 Grosphus voahangyae Lourenço and Wilmé, 2015 Guyanochactas flavus Lourenço and Ythier, 2011 Guyanochactas gonzalezspongai (Lourenço, 1983) Guyanochactas goujei (Vellard, 1932) Guyanochactas mascarenhasi (Lourenço, 1988) Guyanochactas touroulti Lourenço, 2018 Hadogenes austroafricanus Penther, 1900 Hadogenes bicolor Purcell, 1899 Hadogenes gracilis Hewitt, 1909 Hadogenes granulatus Purcell, 1901 Hadogenes gunningi Purcell, 1899 Hadogenes hahni (Peters, 1862) Hadogenes lawrencei Newlands, 1972 Hadogenes longimanus Prendini, 2001 Hadogenes minor Purcell, 1899 Hadogenes newlandsi Prendini, 2001 Hadogenes paucidens Pocock, 1896 Hadogenes phyllodes (Thorell, 1876) Hadogenes polytrichobothrius Prendini, 2006 Hadogenes soutpansbergensis Prendini, 2006 Hadogenes tityrus (Simon, 1888) Hadogenes trichiurus (Gervais, 1843) Hadogenes troglodytes (Peters, 1861) Hadogenes weygoldti Stáhlavsky, Stundlová, Lowe, Stockmann and Kovarík, 2018 Hadogenes zuluanus Lawrence, 1937 Hadogenes zumpti Newlands and Cantrell, 1985 Hadrurochactas araripe Lourenço, 2010 Hadrurochactas brejo (Lourenço, 1988) Hadrurochactas cristinae Ythier, 2018 Hadrurochactas machadoi González-Sponga, 1993 Hadrurochactas odoardoi González-Sponga, 1985 Hadrurochactas polisi (Monod and Lourenço, 2001) Hadrurochactas schaumii (Karsch, 1880) Hadruroides adrianae Rossi, 2012 Hadruroides aguilari Francke and Soleglad, 1980 Hadruroides bustamantei Ochoa and Chaparro, 2008 Hadruroides carinatus Pocock, 1900 Hadruroides charcasus (Karsch, 1879) Hadruroides chinchaysuyu Ochoa and Prendini, 2010 Hadruroides doriai Rossi, 2014 Hadruroides elenae Rossi, 2014 Hadruroides galapagoensis Maury, 1975 Hadruroides geckoi Ochoa and Prendini, 2010 Hadruroides graceae Ochoa and Prendini, 2010 Hadruroides inti Ythier, 2021 Hadruroides juanchaparroi Ochoa and Prendini, 2010 Hadruroides leopardus Pocock, 1900 Hadruroides lourencoi Rossi, 2012 Hadruroides lunatus (L. Koch, 1867) Hadruroides maculatus (Thorell, 1876) Hadruroides mauryi Francke and Soleglad, 1980 Hadruroides moreti Rossi, 2014 Hadruroides pachamama Ythier, 2021 Hadruroides tishqu Ochoa and Prendini, 2010 Hadruroides tongiorgii Rossi, 2012 Hadruroides udvardyi Lourenço, 1995 Hadruroides vichayitos Ochoa and Prendini, 2010 Hadrurus anzaborrego Soleglad, Fet and Lowe, 2011 Hadrurus arizonensis Ewing, 1928 Hadrurus concolorous Stahnke, 1969 Hadrurus hirsutus (Wood, 1863) Hadrurus obscurus Williams, 1970 Hadrurus pinteri Stahnke, 1969 Hadrurus spadix Stahnke, 1940 Hemibuthus crassimanus (Pocock, 1897) Hemibuthus umarii Amir, Kamaluddin and Khan, 2004 Hemilychas alexandrinus (Hirst, 1911) Hemiscorpius acanthocercus Monod and Lourenço, 2005 Hemiscorpius arabicus Pocock, 1899 Hemiscorpius egyptiensis Lourenço, 2011 Hemiscorpius enischnochela Monod and Lourenço, 2005 Hemiscorpius falcifer Lowe, 2010 Hemiscorpius flagelliraptor Lowe, 2010 Hemiscorpius gaillardi (Vachon, 1974) Hemiscorpius kashkayi Karatas and Gharkheloo, 2013 Hemiscorpius lepturus Peters, 1861 Hemiscorpius lipsae Kovarík and Lowe, 2022 Hemiscorpius maindroni (Kraepelin, 1900) Hemiscorpius novaki Kovarík and Mazuch, 2011 Hemiscorpius persicus Birula, 1903 Hemiscorpius shahii Kovarík, Navidpour and Soleglad, 2017 Hemiscorpius socotranus Pocock, 1899 Hemiscorpius somalicus Lourenço, 2011 Hemiscorpius tellinii Borelli, 1904 Heteroctenus abudi (Armas and Marcano Fondeur, 1987) Heteroctenus aridicola (Teruel and Armas, 2012) Heteroctenus bonettii (Armas, 1999) Heteroctenus garridoi (Armas, 1974) Heteroctenus gibarae (Teruel, 2006) Heteroctenus granulimanus (Teruel, 2006) Heteroctenus junceus (Herbst, 1800) Heteroctenus melloleitaoi (Teruel and Armas, 2006) Heteroctenus princeps (Karsch, 1879) Heteroctenus turieli Teruel and Yong, 2023 Heterometrus cimrmani Kovarík, 2004 Heterometrus glaucus (Thorell, 1876) Heterometrus laevigatus (Thorell, 1876) Heterometrus laoticus Couzijn, 1981 Heterometrus liophysa (Thorell, 1888) Heterometrus longimanus (Herbst, 1800) Heterometrus minotaurus Plisková, Kovarík, Kosulic and Stáhlavsky, 2016 Heterometrus petersii (Thorell, 1876) Heterometrus silenus (Simon, 1884) Heterometrus spinifer (Ehrenberg, 1828) Heterometrus thorellii (Pocock, 1892) Heteronebo barahonae Teruel, Armas and Kovarík, 2015 Heteronebo bermudezi (Moreno, 1938) Heteronebo caymanensis Francke, 1978 Heteronebo cicero Armas and Marcano Fondeur, 1987 Heteronebo clareae Armas, 2001 Heteronebo dominicus Armas, 1981 Heteronebo elegans Francke, 1978 Heteronebo forbesii Pocock, 1899 Heteronebo granti Pocock, 1899 Heteronebo jamaicae Francke, 1978 Heteronebo monticola (Armas, 1999) Heteronebo morenoi (Armas, 1973) Heteronebo nibujon Armas, 1984 Heteronebo oviedo (Armas, 1999) Heteronebo portoricensis Francke, 1978 Heteronebo pumilus Armas, 1981 Heteronebo scaber (Pocock, 1893) Heteronebo vachoni Francke, 1978 Heteronebo yntemai Francke and Sissom, 1980 Heteroscorpion goodmani Lourenço, 1996 Heteroscorpion kaii Lourenço and Goodman, 2009 Heteroscorpion kraepelini Lourenço and Goodman, 2006 Heteroscorpion magnus Lourenço and Goodman, 2002 Heteroscorpion opisthacanthoides (Kraepelin, 1896) Heteroscorpion raselimananai Lourenço and Goodman, 2004 Himalayotityobuthus alejandrae Lourenço, 2003 Himalayotityobuthus martensi Lourenço, 1997 Hoffmannihadrurus aztecus (Pocock, 1902) Hoffmannihadrurus gertschi (Soleglad, 1976) Hormiops davidovi Fage, 1933 Hormiops infulcra Monod, 2014 Hormurus boholiensis Kraepelin, 1914 Hormurus ischnoryctes Monod and Prendini in Monod et al., 2013 Hormurus karschii (Keyserling, 1885) Hormurus longimanus (Locket, 1995) Hormurus macrochela Monod and Prendini in Monod et al., 2013 Hormurus neocaledonicus (Simon, 1877) Hormurus ochyroscapter Monod in Monod et al., 2013 Hormurus penta (Francke and Lourenço, 1991) Hormurus polisorum (Volschenk, Locket and Harvey, 2001) Hormurus waigiensis (Gervais, 1843) Hottentotta akbarii Yagmur, Moradi, Tabatabaei and Jafari, 2022 Hottentotta alticola (Pocock, 1895) Hottentotta arenaceus (Purcell, 1901) Hottentotta asimii (Amir, Kamaluddin and Khan, 2004) Hottentotta buchariensis (Birula, 1897) Hottentotta conspersus (Thorell, 1875) Hottentotta finneganae Kovarík, 2007 Hottentotta flavidulus Teruel and Rein, 2010 Hottentotta franzwerneri (Birula, 1914) Hottentotta gambelaensis Kovarík in Kovarík and Mazuch, 2015 Hottentotta gentili (Pallary, 1924) Hottentotta gibaensis Kovarík in Kovarík and Mazuch, 2015 Hottentotta haudensis Kovarík and Lowe, 2021 Hottentotta hoggarensis Lourenço and Leguin, 2014 Hottentotta hottentotta (Fabricius, 1787) Hottentotta jabalpurensis Kovarík, 2007 Hottentotta jalalabadensis Kovarík, 2007 Hottentotta jayakari (Pocock, 1895) Hottentotta judaicus (Simon, 1872) Hottentotta juliae Kovarík, Yagmur and Fet, 2019 Hottentotta keralaensis Aswathi, Sureshan and Lourenço, 2016 Hottentotta khoozestanus Navidpour, Kovarík, Soleglad and Fet, 2008 Hottentotta krivokhatskyi Kovarík, Yagmur and Fet, 2019 Hottentotta lacroixi Ythier and Dupré, 2021 Hottentotta lorestanus Navidpour, Nayebzadeh, Soleglad, Fet, Kovarík and Kayedi, 2010 Hottentotta mazuchi Kovarík in Kovarík and Ojanguren-Affilastro, 2013 Hottentotta mesopotamicus Lourenço and Qi, 2007 Hottentotta minax (L. Koch, 1875) Hottentotta minusalta (Vachon, 1959) Hottentotta navidpouri Kovarík, Yagmur and Moradi, 2018 Hottentotta nigrimontanus Kovarík and Lowe, 2021 Hottentotta novaki Kovarík in Kovarík and Mazuch, 2015 Hottentotta pachyurus (Pocock, 1897) Hottentotta pellucidus Lowe, 2010 Hottentotta penjabensis (Birula, 1897) Hottentotta polystictus (Pocock, 1896) Hottentotta pooyani Moradi, Yagmur, Akbari and Jafari, 2022 Hottentotta reddyi Lourenço, 2015 Hottentotta rugiscutis (Pocock, 1897) Hottentotta salei (Vachon, 1980) Hottentotta saulcyi (Simon, 1880) Hottentotta saxinatans Lowe, 2010 Hottentotta scaber (Ehrenberg, 1828) Hottentotta schach (Birula, 1905) Hottentotta sistanensis Kovarík, Yagmur and Moradi, 2018 Hottentotta socotrensis (Pocock, 1889) Hottentotta somalicus Kovarík, 2018 Hottentotta songi (Lourenço, Qi and Zhu, 2005) Hottentotta sousai Turiel, 2014 Hottentotta stockwelli Kovarík, 2007 Hottentotta syrticus (Borelli, 1914) Hottentotta tamulus (Fabricius, 1798) Hottentotta trailini Kovarík in Kovarík and Ojanguren-Affilastro, 2013 Hottentotta trilineatus (Peters, 1861) Hottentotta ugandaensis Kovarík in Kovarík and Ojanguren-Affilastro, 2013 Hottentotta vinchu Mirza, Ambekar and Kulkarni, 2019 Hottentotta zagrosensis Kovarík, 1997 Iomachus ineichi Lourenço, 2020 Iomachus laeviceps (Pocock, 1890) Iomachus malabarensis Pocock, 1900 Iomachus nitidus Pocock, 1900 Iomachus politus Pocock, 1896 Iomachus punctulatus Pocock, 1897 Iomachus surgani Bastawade, 1986 Iranobuthus krali Kovarík, 1997 Ischnotelson guanambiensis (Lenarducci, Pinto-da-Rocha and Lucas, 2005) Ischnotelson peruassu Esposito, Yamaguti, Souza, Pinto-da-Rocha and Prendini, 2017 Isometroides vescus (Karsch, 1880) Isometrus amboli Sulakhe, Dandekar, Padhye and Bastawade, 2020 Isometrus atherii Amir and Kamaluddin, 2008 Isometrus formosus Pocock, 1894 Isometrus isadensis Tikader and Bastawade, 1983 Isometrus kovariki Sulakhe, Dandekar, Mukherjee, Pandey, Ketkar, Padhye and Bastawade, 2020 Isometrus lao Lourenço and Leguin, 2012 Isometrus liaqatii Amir and Kamaluddin, 2008 Isometrus longitelson Deshpande, Gowande, Bastawade and Sulakhe, 2022 Isometrus maculatus (De Geer, 1778) Isometrus nakshatra Sulakhe, Deshpande, Gowande, Dandekar and Ketkar, 2022 Isometrus sankeriensis Tikader and Bastawade, 1983 Isometrus tamhini Sulakhe, Dandekar, Padhye and Bastawade, 2020 Isometrus thurstoni Pocock, 1893 Isometrus thwaitesi Pocock, 1897 Isometrus wayanadensis Sulakhe, Deshpande, Gowande, Dandekar and Ketkar, 2022 Iurus dekanum (Roewer, 1943) Iurus dufoureius (Brullé, 1832) Iurus kinzelbachi Kovarík, Fet, Soleglad and Yagmur, 2010 Jaguajir agamemnon (C. L. Koch, 1839) Jaguajir pintoi (Mello-Leitão, 1932) Jaguajir rochae (Borelli, 1910) Janalychas albimanus (Henderson, 1919) Janalychas farkasi (Kovarík, 1997) Janalychas granulatus Mirza, 2020 Janalychas heurtaultae (Kovarík, 1997) Janalychas keralaensis Mirza, 2020 Janalychas laevifrons (Pocock, 1897) Janalychas shoplandi (Oates, 1888) Janalychas srilankensis (Lourenço, 1997) Janalychas tricarinatus (Simon, 1884) Javanimetrus cyaneus (C. L. Koch, 1836) Karasbergia methueni Hewitt, 1913 Kochius barbatus (Williams, 1971) Kochius bruneus (Williams, 1970) Kochius cerralvensis (Williams, 1971) Kochius colluvius Ayrey, Jones and Myers, 2019 Kochius hirsuticauda (Banks, 1910) Kochius insularis (Williams, 1971) Kochius magdalensis (Williams, 1971) Kochius punctipalpi (Wood, 1863) Kochius sonorae (Williams, 1971) Kochius villosus (Williams, 1971) Kolotl magnus (Beutelspacher and López-Forment, 1991) Kolotl poncei (Francke and Quijano-Ravell, 2009) Konetontli acapulco (Armas and Martín-Frías, 2001) Konetontli chamelaensis (Williams, 1986) Konetontli ignes González-Santillán and Prendini, 2015 Konetontli ilitchi González-Santillán and Prendini, 2015 Konetontli juxtlahuaca González-Santillán and Prendini, 2015 Konetontli kuarapu (Francke and Ponce Saavedra, 2005) Konetontli migrus González-Santillán and Prendini, 2015 Konetontli nayarit (Armas and Martín-Frías, 2001) Konetontli pattersoni (Williams and Haradon in Williams, 1980) Konetontli zihuatanejensis (Baldazo-Monsivaiz, 2003) Kovarikia angelena (Gertsch and Soleglad, 1972) Kovarikia bogerti (Gertsch and Soleglad, 1972) Kovarikia oxy Bryson, Graham and Soleglad in Bryson et al., 2018 Kovarikia savaryi Bryson, Graham and Soleglad in Bryson et al., 2018 Kovarikia williamsi (Gertsch and Soleglad, 1972) Kraepelinia palpator (Birula, 1903) Kuarapu purhepecha Francke and Ponce Saavedra, 2010 Langxie feti Tang, Jia and Liu, 2023 Lanzatus huluul Kovarík and Lowe, 2021 Lanzatus somalicus Kovarík, 2001 Lanzatus somalilandus Kovarík and Lowe in Kovarík et al., 2016 Leiurus abdullahbayrami Yagmur, Koc and Kunt, 2009 Leiurus aegyptiacus Lourenço and El-Hennawy, 2021 Leiurus arabicus Lowe, Yagmur and Kovarík, 2014 Leiurus ater Lourenço, 2019 Leiurus brachycentrus (Ehrenberg, 1829) Leiurus dekeyseri Lourenço, 2020 Leiurus gubanensis Kovarík and Lowe, 2020 Leiurus haenggii Lowe, Yagmur and Kovarík, 2014 Leiurus heberti Lowe, Yagmur and Kovarík, 2014 Leiurus hebraeus (Birula, 1908) Leiurus hoggarensis Lourenço, Kourim and Sadine, 2018 Leiurus jordanensis Lourenço, Modry and Amr, 2002 Leiurus kuwaiti Lourenço, 2020 Leiurus macroctenus Lowe, Yagmur and Kovarík, 2014 Leiurus maculatus Lourenço, 2022 Leiurus nigerianus Lourenço, 2021 Leiurus quinquestriatus (Ehrenberg, 1828) Leiurus saharicus Lourenço, 2020 Leiurus savanicola Lourenço, Qi and Cloudsley-Thompson, 2006 Leiurus somalicus Lourenço and Rossi, 2016 Letoiurus rhodiensis (Soleglad, Fet, Kovarík and Yagmur, 2012) Liobuthus kessleri Birula, 1898 Liocheles australasiae (Fabricius, 1775) Liocheles boholiensis (Kraepelin, 1914) Liocheles litodactylus Monod and Volschenk, 2004 Liocheles longimanus (Werner, 1939) Liocheles neocaledonicus (Simon, 1877) Liocheles nigripes (Pocock, 1897) Liocheles oranghutan Ythier and Richard, 2020 Liocheles schalleri Mirza, 2017 Lisposoma elegans Lawrence, 1928 Lisposoma haringtoni (Prendini, 2003) Lisposoma josehermana Lamoral, 1979 Lissothus bernardi Vachon, 1948 Lissothus chaambi Lourenço and Sadine, 2014 Lissothus occidentalis Vachon, 1950 Lychas aberlenci Lourenço, 2013 Lychas armasi Kovarík, 2013 Lychas armillatus (Gervais, 1841) Lychas asper (Pocock, 1891) Lychas biharensis Tikader and Bastawade, 1983 Lychas brehieri Lourenço, 2017 Lychas buchari Kovarík, 1997 Lychas cernickai Kovarík, 2013 Lychas chanthaburiensis Ythier and Lourenço, 2022 Lychas flavimanus (Thorell, 1888) Lychas gravelyi Henderson, 1913 Lychas hendersoni (Pocock, 1897) Lychas hillyardi Kovarík, 1997 Lychas inexpectatus Lourenço, 2011 Lychas jakli Kovarík, 2023 Lychas kamshetensis Tikader and Bastawade, 1983 Lychas kharpadi Bastawade, 1986 Lychas krali Kovarík, 1995 Lychas lourencoi Kovarík, 1997 Lychas marmoreus (C. L. Koch, 1844) Lychas mjobergi Kraepelin, 1916 Lychas mucronatus (Fabricius, 1798) Lychas nigristernis (Pocock, 1899) Lychas obsti Kraepelin, 1913 Lychas perfidus (Keyserling, 1885) Lychas rackae Kovarík, 1997 Lychas rugosus (Pocock, 1897) Lychas santoensis Lourenço, 2009 Lychas scaber (Pocock, 1893) Lychas scutilus C. L. Koch, 1845 Lychas serratus (Pocock, 1891) Lychas shelfordi (Borelli, 1904) Lychas variatus (Thorell, 1876) Lychasioides amieti Vachon, 1974 Maaykuyak vittatus (Williams, 1970) Maaykuyak waueri (Gertsch and Soleglad, 1972) Mauritanobuthus geniezi Qi and Lourenço, 2007 Mauryius cuyanus Ojanguren-Affilastro and Mattoni, 2017 Megachactops coriacea (González-Sponga, 1991) Megachactops kuemoi Ochoa, Rojas-Runjaic, Pinto-da-Rocha and Prendini, 2013 Megachactops kurripako Ythier, 2019 Megacormus franckei Kovarík, 2019 Megacormus gertschi Díaz Najera, 1966 Megacormus granosus (Gervais, 1843) Megacormus grubbsi Sissom, 1994 Megacormus orizaba Teruel, Kovarík, Lowe and Stáhlavský, 2023 Megacormus seductus Teruel, Kovarík, Lowe and Stáhlavský, 2023 Megacormus segmentatus Pocock, 1900 Megacormus xichu González-Santillán, González-Ruíz and Escobedo-Morales, 2017 Mesobuthus afghanus (Pocock, 1889) Mesobuthus barszczevskii (Birula, 1904) Mesobuthus birulai Kovarík et al., 2022 Mesobuthus bogdoensis (Birula, 1896) Mesobuthus crucittii Kovarík et al., 2022 Mesobuthus eupeus (C. L. Koch, 1839) Mesobuthus farleyi Kovarík et al., 2022 Mesobuthus fomichevi Kovarík et al., 2022 Mesobuthus galinae Kovarík et al., 2022 Mesobuthus haarlovi Vachon, 1958 Mesobuthus iranus (Birula, 1917) Mesobuthus kaftani Kovarík et al., 2022 Mesobuthus kirmanensis (Birula, 1900) Mesobuthus macmahoni (Pocock, 1900) Mesobuthus marusiki Kovarík et al., 2022 Mesobuthus mesopotamicus (Penther, 1912) Mesobuthus mirshamsii Kovarík et al., 2022 Mesobuthus navidpouri Kovarík et al., 2022 Mesobuthus persicus (Pocock, 1899) Mesobuthus philippovitschi (Birula, 1905) Mesobuthus phillipsii (Pocock, 1889) Mesobuthus rahsenae Kovarík et al., 2022 Mesobuthus rakhshanii Barahoei, 2022 Mesobuthus thersites (C. L. Koch, 1839) Mesobuthus turcicus Kovarík et al., 2022 Mesobuthus vesiculatus (Pocock, 1899) Mesobuthus vignolii Kovarík et al., 2022 Mesobuthus yagmuri Kovarík and Fet in Kovarík et al., 2022 Mesobuthus zarudnyi Novruzov, Kovarík and Fet, 2022 Mesobuthus zonsteini Kovarík et al., 2022 Mesomexovis atenango (Francke and González-Santillán, 2007) Mesomexovis oaxaca (Santibañez-López and Sissom, 2010) Mesomexovis occidentalis (Hoffmann, 1931) Mesomexovis punctatus (Karsch, 1879) Mesomexovis spadix (Hoffmann, 1931) Mesomexovis subcristatus (Pocock, 1898) Mesomexovis variegatus (Pocock, 1898) Mesotityus vondangeli González-Sponga, 1981 Metaiurus kadleci (Kovarík, Fet, Soleglad and Yagmur, 2010) Metaiurus stathiae (Soleglad, Fet, Kovarík and Yagmur, 2012) Microananteris abounami Lourenço and Chevalier, 2022 Microananteris inselberg Lourenço, 2021 Microananteris minor Lourenço, 2003 Microananteris serrulata Lourenço, 2021 Microbuthus fagei Vachon, 1949 Microbuthus flavorufus Lourenço and Duhem, 2007 Microbuthus gardneri Lowe, 2010 Microbuthus kristensenorum Lowe, 2010 Microbuthus litoralis (Pavesi, 1885) Microbuthus maroccanus Lourenço, 2002 Microbuthus satyrus Lowe, Kovarík, Stockmann and Stáhlavsky, 2018 Microcharmus andrei Lourenço, Waeber and Wilmé, 2019 Microcharmus antongil Lourenço, Waeber and Wilmé, 2019 Microcharmus bemaraha Lourenço, Goodman and Fisher, 2006 Microcharmus cloudsleythompsoni Lourenço, 1995 Microcharmus confluenciatus Lourenço, Goodman and Fisher, 2006 Microcharmus djangoa Lourenço, Waeber and Wilmé, 2019 Microcharmus duhemi Lourenço, Goodman and Fisher, 2006 Microcharmus fisheri Lourenço, 1998 Microcharmus hauseri Lourenço, 1996 Microcharmus henderickxi Lourenço, 2009 Microcharmus jussarae Lourenço, 1996 Microcharmus maculatus Lourenço, Goodman and Fisher, 2006 Microcharmus madagascariensis Lourenço and Goodman, 1999 Microcharmus pauliani (Lourenço, 2004) Microcharmus sabineae Lourenço, 1996 Microcharmus variegatus Lourenço, Goodman and Fisher, 2006 Microcharmus violaceus Lourenço, Goodman and Fisher, 2006 Microtityus ambarensis (Schawaller, 1981) Microtityus angelaerrosae González-Sponga, 2001 Microtityus barahona Armas and Teruel, 2012 Microtityus biordi González-Sponga, 1970 Microtityus bivicentorum Botero-Trujillo, Erazo-Moreno and Pérez, 2009 Microtityus borincanus Teruel, Rivera and Sánchez, 2014 Microtityus capayaensis González-Sponga, 2001 Microtityus consuelo Armas and Marcano Fondeur, 1987 Microtityus desuzeae González-Sponga, 2001 Microtityus difficilis Teruel and Armas, 2006 Microtityus dominicanensis Santiago-Blay, 1985 Microtityus eustatia Armas, 2018 Microtityus farleyi Teruel, 2000 Microtityus flavescens Teruel, 2001 Microtityus franckei Botero-Trujillo and Noriega, 2008 Microtityus fundorai Armas, 1974 Microtityus guantanamo Armas, 1984 Microtityus iviei Armas, 1999 Microtityus jaumei Armas, 1974 Microtityus joseantonioi González-Sponga, 1981 Microtityus kovariki Teruel and Infante, 2007 Microtityus lantiguai Armas and Marcano Fondeur, 1992 Microtityus litoralensis González-Sponga, 2001 Microtityus lourencoi Armas and Teruel, 2012 Microtityus minimus Kovarík and Teruel, 2014 Microtityus paucidentatus Armas and Marcano Fondeur, 1992 Microtityus prendinii Armas and Teruel, 2012 Microtityus pusillus Teruel and Kovarík, 2012 Microtityus reini Armas and Teruel, 2012 Microtityus rickyi Kjellesvig-Waering, 1966 Microtityus santosi Teruel, Rivera and Sánchez, 2014 Microtityus sevciki González-Sponga, 2001 Microtityus solegladi Armas and Teruel, 2012 Microtityus starri Lourenço and Huber, 1999 Microtityus trinitensis Armas, 1974 Microtityus vanzolinii Lourenço and Eickstedt, 1983 Microtityus vieques Teruel, Rivera and Santos, 2015 Microtityus virginiae Armas, 1999 Microtityus vulcanicus Teruel, 2019 Microtityus waeringi Francke and Sissom, 1980 Microtityus yaracuyanus González-Sponga, 2001 Mioscorpio zeuneri (Hadzi, 1931) Nebo flavipes Simon, 1882 Nebo franckei Vachon, 1980 Nebo grandis Francke, 1980 Nebo henjamicus Francke, 1980 Nebo hierichonticus (Simon, 1872) Nebo jazanensis Abu Afifeh, Aloufi, Al-Saraireh and Amr, 2023 Nebo omanensis Francke, 1980 Nebo poggesii Sissom, 1994 Nebo whitei Vachon, 1980 Nebo yemenensis Francke, 1980 Neobuthus amoudensis Kovarík, Lowe, Awale, Elmi and Hurre, 2018 Neobuthus awashensis Kovarík and Lowe, 2012 Neobuthus berberensis Hirst, 1911 Neobuthus cloudsleythompsoni Lourenço, 2001 Neobuthus erigavoensis Kovarík, Lowe, Awale, Elmi and Hurre, 2018 Neobuthus eritreaensis Lowe and Kovarík, 2016 Neobuthus factorio Kovarík, Lowe, Awale, Elmi and Hurre, 2018 Neobuthus ferrugineus (Kraepelin, 1898) Neobuthus gubanensis Kovarík, Lowe, Awale, Elmi and Hurre, 2018 Neobuthus haeckeli Kovarík, 2019 Neobuthus kloppersi Kovarík, Lowe, Awale, Elmi and Hurre, 2018 Neobuthus kutcheri Lowe and Kovarík, 2016 Neobuthus maidensis Kovarík, Lowe, Awale, Elmi and Hurre, 2018 Neobuthus montanus Kovarík, Lowe, Awale, Elmi and Hurre, 2018 Neobuthus solegladi Kovarík, 2019 Neobuthus sudanensis Lourenço, 2005 Neocalchas gruberi (Fet, Soleglad and Kovarík, 2009) Neochactas bariensis (González-Sponga, 1991) Neochactas bilbaoi (González-Sponga, 1978) Neochactas bruzuali (González-Sponga, 1980) Neochactas caroniensis (González-Sponga, 1996) Neochactas colombiensis (González-Sponga, 1976) Neochactas delicatus (Karsch, 1879) Neochactas efreni (González-Sponga, 1978) Neochactas eliasilvai (González-Sponga, 1980) Neochactas fei (Pinto-da-Rocha and Brescovit in Pinto-da-Rocha et al., 2002) Neochactas fravalae (Lourenço, 1983) Neochactas gaillardi (Lourenço, 1983) Neochactas garciai (González-Sponga, 1978) Neochactas granosus (Pocock, 1900) Neochactas guaiquinimensis (González-Sponga, 1997) Neochactas jaspei (González-Sponga, 1993) Neochactas josemanueli (González-Sponga, 1992) Neochactas kelleri (Lourenço, 1997) Neochactas kjellesvigi (González-Sponga, 1974) Neochactas laui (Kjellesvig-Waering, 1966) Neochactas leoneli (González-Sponga, 1978) Neochactas macrochelae (González-Sponga, 2004) Neochactas mottai (Lourenço and Araujo, 2004) Neochactas neblinensis (González-Sponga, 1991) Neochactas orinocensis (Scorza, 1954) Neochactas panarei (González-Sponga, 1980) Neochactas paoensis (González-Sponga, 1996) Neochactas parvulus (Pocock, 1897) Neochactas racenisi (González-Sponga, 1975) Neochactas riopinensis (González-Sponga, 1992) Neochactas ruizpittoli (González-Sponga, 1993) Neochactas sanmartini (González-Sponga, 1974) Neochactas santanai (González-Sponga, 1978) Neochactas sarisarinamensis (González-Sponga, 1985) Neochactas simarawochensis (González-Sponga, 1980) Neochactas sissomi (Lourenço, 1983) Neochactas skuki (Lourenço and Pinto-da-Rocha, 2000) Neochactas verai (González-Sponga, 1993) Neochactas yekuanae (González-Sponga, 1984) Neogrosphus andrafiabe Lourenço, Wilmé and Waeber, 2015 Neogrosphus blanci Lourenço, 1996 Neogrosphus griveaudi (Vachon, 1969) Neoprotobuthus intermedius Lourenço, 2000 Nullibrotheas allenii (Wood, 1863) Odontobuthus baluchicus Barahoei et al., 2021 Odontobuthus bidentatus Lourenço and Pézier, 2002 Odontobuthus brevidigitus Lowe, 2010 Odontobuthus chabaharensis Barahoei et al., 2021 Odontobuthus doriae (Thorell, 1876) Odontobuthus kermanus Barahoei et al., 2021 Odontobuthus odonturus (Pocock, 1897) Odontobuthus tavighiae Navidpour, Soleglad, Fet and Kovarík, 2013 Odontobuthus tirgari Mirshamsi, Azghadi, Navidpour, Aliabadian and Kovarík, 2013 Odonturus dentatus Karsch, 1879 Oiclus ardens Ythier, 2019 Oiclus cousteaui Ythier, 2019 Oiclus nanus Teruel and Chazal, 2010 Oiclus purvesii (Becker, 1880) Oiclus questeli Teruel, 2008 Oiclus tipunch Ythier, 2019 Oiclus tite Ythier, Jourdan and Malglaive, 2022 Olivierus bolensis (Sun, Zhu and Lourenço, 2010) Olivierus brutus (Fet, Kovarík, Gantenbein, Kaiser, Stewart and Graham, 2018) Olivierus caucasicus (Nordmann, 1840) Olivierus elenae (Fet, Kovarík, Gantenbein, Kaiser, Stewart and Graham, 2018) Olivierus extremus (Werner, 1936) Olivierus fuscus (Birula, 1897) Olivierus gorelovi (Fet, Kovarík, Gantenbein, Kaiser, Stewart and Graham, 2018) Olivierus hainanensis (Birula, 1904) Olivierus intermedius (Birula, 1897) Olivierus karshius (Sun and Z.-N. Sun, 2011) Olivierus kaznakovi (Birula, 1904) Olivierus kreuzbergi (Fet, Kovarík, Gantenbein, Kaiser, Stewart and Graham, 2018) Olivierus longichelus (Sun and Zhu, 2010) Olivierus martensii (Karsch, 1879) Olivierus mikhailovi Fet, Kovarík, Gantenbein and Graham, 2021 Olivierus mischi (Fet, Kovarík, Gantenbein, Kaiser, Stewart and Graham, 2018) Olivierus nenilini (Fet, Kovarík, Gantenbein, Kaiser, Stewart and Graham, 2018) Olivierus parthorum (Pocock, 1889) Olivierus przewalskii (Birula, 1897) Olivierus tarabaevi Fet, Kovarík, Gantenbein and Graham, 2021 Olivierus voldemari Fet, Kovarík, Gantenbein and Graham, 2021 Opisthacanthus africanus Simon, 1876 Opisthacanthus ambanja Lourenço, 2014 Opisthacanthus andohahela Lourenço, 2014 Opisthacanthus antsiranana Lourenço, 2014 Opisthacanthus asper (Peters, 1861) Opisthacanthus autanensis González-Sponga, 2006 Opisthacanthus basutus Lawrence, 1955 Opisthacanthus borboremai Lourenço and Fé, 2003 Opisthacanthus brevicauda Rojas-Runjaic, Borges and Armas, 2008 Opisthacanthus capensis Thorell, 1876 Opisthacanthus cayaporum Vellard, 1932 Opisthacanthus darainensis Lourenço and Goodman, 2006 Opisthacanthus diremptus (Karsch, 1879) Opisthacanthus elatus (Gervais, 1844) Opisthacanthus faillei Lourenço and Wilmé, 2019 Opisthacanthus heurtaultae Lourenço, 1980 Opisthacanthus laevipes (Pocock, 1893) Opisthacanthus lamorali Lourenço, 1981 Opisthacanthus lavasoa Lourenço, Wilmé and Waeber, 2016 Opisthacanthus lecomtei (Lucas, 1858) Opisthacanthus lepturus (Palisot de Beauvois, 1805) Opisthacanthus lourencoi Ythier, 2022 Opisthacanthus lucienneae Lourenço and Goodman, 2006 Opisthacanthus maculatus Lourenço and Goodman, 2006 Opisthacanthus madagascariensis Kraepelin, 1894 Opisthacanthus milloti Lourenço and Goodman, 2008 Opisthacanthus pauliani Lourenço and Goodman, 2008 Opisthacanthus piceus Lourenço and Goodman, 2006 Opisthacanthus piscatorius Lawrence, 1955 Opisthacanthus punctulatus Pocock, 1896 Opisthacanthus rugiceps Pocock, 1897 Opisthacanthus rugulosus Pocock, 1896 Opisthacanthus surinamensis Lourenço, 2017 Opisthacanthus titanus Lourenço, Wilmé and Waeber, 2018 Opisthacanthus valerioi Lourenço, 1980 Opisthacanthus validus Thorell, 1876 Opisthacanthus weyrauchi Mello-Leitão, 1948 Opistophthalmus adustus Kraepelin, 1908 Opistophthalmus ammopus Lamoral, 1980 Opistophthalmus ater Purcell, 1898 Opistophthalmus austerus Karsch, 1879 Opistophthalmus boehmi (Kraepelin, 1896) Opistophthalmus brevicauda Lawrence, 1928 Opistophthalmus capensis (Herbst, 1800) Opistophthalmus carinatus (Peters, 1861) Opistophthalmus cavimanus Lawrence, 1928 Opistophthalmus chaperi Simon, 1880 Opistophthalmus chrysites Lawrence, 1967 Opistophthalmus coetzeei Lamoral, 1979 Opistophthalmus concinnus Newlands, 1972 Opistophthalmus crassimanus Purcell, 1899 Opistophthalmus fitzsimonsi Hewitt, 1935 Opistophthalmus flavescens Purcell, 1898 Opistophthalmus fossor Purcell, 1898 Opistophthalmus fuscipes Purcell, 1898 Opistophthalmus gibbericauda Lamoral, 1979 Opistophthalmus gigas Purcell, 1898 Opistophthalmus glabrifrons Peters, 1861 Opistophthalmus granicauda Purcell, 1898 Opistophthalmus granifrons Pocock, 1896 Opistophthalmus haackei Lawrence, 1966 Opistophthalmus harpei Harington, 2002 Opistophthalmus holmi (Lawrence, 1969) Opistophthalmus intercedens Kraepelin, 1908 Opistophthalmus intermedius Kraepelin, 1894 Opistophthalmus jenseni (Lamoral, 1972) Opistophthalmus karrooensis Purcell, 1898 Opistophthalmus keilandsi Hewitt, 1914 Opistophthalmus lamorali Prendini, 2000 Opistophthalmus laticauda Purcell, 1898 Opistophthalmus latimanus C. L. Koch, 1841 Opistophthalmus latro Thorell, 1876 Opistophthalmus lawrencei Newlands, 1969 Opistophthalmus leipoldti Purcell, 1898 Opistophthalmus litoralis Lawrence, 1955 Opistophthalmus longicauda Purcell, 1899 Opistophthalmus lornae Lamoral, 1979 Opistophthalmus luciranus Lawrence, 1959 Opistophthalmus macer Thorell, 1876 Opistophthalmus nitidiceps Pocock, 1896 Opistophthalmus opinatus (Simon, 1888) Opistophthalmus pallipes C. L. Koch, 1842 Opistophthalmus pattisoni Purcell, 1899 Opistophthalmus penrithorum Lamoral, 1979 Opistophthalmus peringueyi Purcell, 1898 Opistophthalmus pictus Kraepelin, 1894 Opistophthalmus pluridens Hewitt, 1918 Opistophthalmus praedo Thorell, 1876 Opistophthalmus pugnax Thorell, 1876 Opistophthalmus pygmaeus Lamoral, 1979 Opistophthalmus scabrifrons Hewitt, 1918 Opistophthalmus schlechteri Purcell, 1898 Opistophthalmus schultzei Kraepelin, 1908 Opistophthalmus setifrons Lawrence, 1961 Opistophthalmus tumas Ythier, 2023 Opistophthalmus ugabensis Hewitt, 1934 Opistophthalmus wahlbergii (Thorell, 1876) Orobothriurus alticola (Pocock, 1899) Orobothriurus ampay Ochoa and Acosta, 2003 Orobothriurus atiquipa Ochoa and Acosta, 2002 Orobothriurus calchaqui Ochoa, Ojanguren-Affilastro, Mattoni and Prendini, 2011 Orobothriurus compagnucii Ochoa, Ojanguren-Affilastro, Mattoni and Prendini, 2011 Orobothriurus curvidigitus (Kraepelin, 1911) Orobothriurus famatina Acosta, 2001 Orobothriurus grismadoi Ojanguren-Affilastro, Fernández Campón, Silnik and Mattoni, 2009 Orobothriurus huascaran Ochoa, Ojanguren-Affilastro, Mattoni and Prendini, 2011 Orobothriurus paessleri (Kraepelin, 1911) Orobothriurus parvus Maury, 1975 Orobothriurus quewerukana Ochoa, Ojanguren-Affilastro, Mattoni and Prendini, 2011 Orobothriurus ramirezi Ochoa, Ojanguren-Affilastro, Mattoni and Prendini, 2011 Orobothriurus tamarugal Ochoa, Ojanguren-Affilastro, Mattoni and Prendini, 2011 Orobothriurus wawita Acosta and Ochoa, 2000 Orthochiroides insularis (Pocock, 1899) Orthochiroides socotrensis Kovarík, 2004 Orthochiroides somalilandus Kovarík and Lowe, 2022 Orthochiroides vachoni Kovarík, 1998 Orthochirus afar Kovarík and Lowe in Kovarík et al., 2016 Orthochirus afghanus Kovarík, 2004 Orthochirus arabicus Ythier and Lourenço, 2023 Orthochirus aristidis (Simon, 1882) Orthochirus atarensis Lourenço and Leguin, 2011 Orthochirus bastawadei Zambre, Mirza, Sanap, Upadhye and Javed, 2011 Orthochirus bicolor (Pocock, 1897) Orthochirus birulai Kovarík, Fet and Yagmur, 2020 Orthochirus carinatus Navidpour, Kovarík, Soleglad and Fet, 2019 Orthochirus cloudsleythompsoni Lourenço and Leguin, 2011 Orthochirus danielleae (Lourenço and Vachon, 1997) Orthochirus farzanpayi (Vachon and Farzanpay, 1987) Orthochirus feti Kovarík, 2004 Orthochirus flavescens (Pocock, 1897) Orthochirus fomichevi Kovarík, Yagmur, Fet and Hussen, 2019 Orthochirus formozovi Kovarík, Fet and Yagmur, 2020 Orthochirus fuscipes (Pocock, 1900) Orthochirus gantenbeini Kovarík, Yagmur, Fet and Hussen, 2019 Orthochirus glabrifrons (Kraepelin, 1903) Orthochirus gromovi Kovarík, 2004 Orthochirus grosseri Kovarík, Fet and Yagmur, 2020 Orthochirus gruberi Kovarík and Fet, 2006 Orthochirus heratensis Kovarík, 2004 Orthochirus hormozganensis Kovarík and Navidpour, 2020 Orthochirus innesi Simon, 1910 Orthochirus iranus Kovarík, 2004 Orthochirus iraqus Kovarík, 2004 Orthochirus jalalabadensis Kovarík, 2004 Orthochirus kaspareki (Lourenço and Huber, 2000) Orthochirus katerinae Kovarík and Just, 2022 Orthochirus kermanensis Kovarík and Navidpour, 2020 Orthochirus kinzelbachi (Lourenço and Huber, 2000) Orthochirus kovariki Yagmur and Khalili, 2022 Orthochirus krishnai Tikader and Bastawade, 1983 Orthochirus kryzhanovskyi Kovarík, Fet and Yagmur, 2020 Orthochirus kucerai Kovarík and Navidpour, 2020 Orthochirus maroccanus Lourenço and Leguin, 2011 Orthochirus masihipouri Kovarík and Navidpour, 2020 Orthochirus melanurus (Kessler, 1874) Orthochirus mesopotamicus Birula, 1918 Orthochirus milloti Lourenço, 2021 Orthochirus minor Lourenço, Duhem and Cloudsley-Thompson, 2012 Orthochirus monodi (Lourenço and Vachon, 1997) Orthochirus navidpouri Kovarík, Yagmur, Fet and Hussen, 2019 Orthochirus nordmanni Kovarík, Fet and Yagmur, 2020 Orthochirus olivaceus (Karsch, 1881) Orthochirus pallidus (Pocock, 1897) Orthochirus persa (Birula, 1900) Orthochirus samrchelsis Kovarík, 2004 Orthochirus scrobiculosus (Grube, 1873) Orthochirus sejnai Kovarík, Fet and Yagmur, 2020 Orthochirus semnanensis Kovarík and Navidpour, 2020 Orthochirus soufiensis Lourenço and Sadine, 2021 Orthochirus stockwelli (Lourenço and Vachon, 1995) Orthochirus tassili Lourenço and Leguin, 2011 Orthochirus tibesti Lourenço, Duhem and Cloudsley-Thompson, 2012 Orthochirus varius Kovarík, 2004 Orthochirus vignolii Kovarík and Navidpour, 2020 Orthochirus zagrosensis Kovarík, 2004 Pachakutej crassimanus (Maury, 1975) Pachakutej inca (Maury, 1975) Pachakutej iskay (Acosta and Ochoa, 2001) Pachakutej juchuicha Ochoa, 2004 Pachakutej oscari Ochoa, 2004 Pachakutej peruvianus (Mello-Leitão, 1948) Palaeoakentrobuthus knodeli Lourenço and Weitschat, 2000 Palaeoananteris ribnitiodamgartensis Lourenço and Weitschat, 2001 Palaeoananteris ukrainensis Lourenço and Weitschat, 2009 Palaeoananteris wunderlichi Lourenço, 2004 Palaeoburmesebuthus grimaldii Lourenço, 2002 Palaeoburmesebuthus longipalpis Lourenço and Rossi, 2017 Palaeoburmesebuthus ohlhoffi Lourenço, 2015 Palaeocheloctonus pauliani Lourenço, 1996 Palaeocheloctonus septentrionalis Lourenço and Wilmé, 2015 Palaeoeuscorpius gallicus Lourenço, 2003 Palaeogrosphus copalensis (Lourenço, 1996) Palaeogrosphus jacquesi Lourenço and Henderickx, 2012 Palaeoisometrus elegans Lourenço and Weitschat, 2005 Palaeolychas balticus Lourenço and Weitschat, 1996 Palaeolychas weitschati Lourenço, 2012 Palaeoprotobuthus pusillus Lourenço and Weitschat, 2000 Palaeospinobuthus cenozoicus Lourenço, Henderickx and Weitschat, 2005 Palaeotityobuthus longiaculeus Lourenço and Weitschat, 2000 Palaeotrilineatus ellenbergeri Lourenço, 2012 Pandiborellius arabicus (Kraepelin, 1894) Pandiborellius awashensis (Kovarík, 2012) Pandiborellius igdu Kovarík, Lowe, Soleglad and Plísková, 2017 Pandiborellius insularis Kovarík, Lowe, Soleglad and Plísková, 2017 Pandiborellius lanzai (Rossi, 2015) Pandiborellius magretti (Borelli, 1901) Pandiborellius meidensis (Karsch, 1879) Pandiborellius nistriae (Rossi, 2014) Pandiborellius percivali (Pocock, 1902) Pandiborellius somalilandus (Kovarík, 2012) Pandinoides cavimanus (Pocock, 1888) Pandinoides duffmackayi Prendini, 2016 Pandinoides militaris (Pocock, 1900) Pandinops bambii Rossi, 2016 Pandinops bellicosus (L. Koch, 1875) Pandinops colei (Pocock, 1896) Pandinops eritreaensis (Kovarík, 2003) Pandinops friedrichi Kovarík, 2016 Pandinops pococki (Kovarík, 2000) Pandinops pugilator (Pocock, 1900) Pandinops turieli Kovarík, 2016 Pandinopsis dictator (Pocock, 1888) Pandinurus afar Kovarík, Lowe, Soleglad and Plísková, 2017 Pandinurus awalei Kovarík, Lowe and Elmi, 2020 Pandinurus citernii (Borelli, 1919) Pandinurus exitialis (Pocock, 1888) Pandinurus fulvipes Kovarík, Lowe and Mazuch, 2019 Pandinurus gregoryi (Pocock, 1896) Pandinurus hangarale Kovarík, Lowe, Mazuch, Awale, Stundlová and Stáhlavsky, 2017 Pandinurus intermedius (Borelli, 1919) Pandinurus kmoniceki Kovarík, Lowe, Mazuch, Plísková and Stáhlavsky, 2017 Pandinurus mazuchi (Kovarík, 2011) Pandinurus oromo Kovarík, Lowe, Soleglad and Plísková, 2017 Pandinurus pallidus (Kraepelin, 1894) Pandinurus phillipsii (Pocock, 1896) Pandinurus platycheles (Werner, 1916) Pandinurus sabbadinii Rossi, 2015 Pandinurus smithi (Pocock, 1899) Pandinurus sudanicus (Hirst, 1911) Pandinurus trailini (Kovarík, 2013) Pandinus boschisi Caporiacco, 1937 Pandinus gambiensis Pocock, 1899 Pandinus imperator (C. L. Koch, 1841) Pandinus ugandaensis Kovarík, 2011 Pandinus ulderigoi Rossi, 2014 Pandipalpus lowei (Kovarík, 2012) Pandipalpus viatoris (Pocock, 1890) Pantobuthus complicatus Lourenço and Duhem, 2009 Parabuthus abyssinicus Pocock, 1901 Parabuthus brevimanus (Thorell, 1876) Parabuthus calvus Purcell, 1898 Parabuthus capensis (Ehrenberg, 1831) Parabuthus cimrmani Kovarík, 2004 Parabuthus distridor Lamoral, 1980 Parabuthus erigavoensis Kovarík, Lowe, Elmi and Stáhlavsky, 2019 Parabuthus eritreaensis Kovarík, 2003 Parabuthus glabrimanus Prendini and Esposito, 2010 Parabuthus gracilis Lamoral, 1979 Parabuthus granimanus Pocock, 1895 Parabuthus granulatus (Ehrenberg, 1831) Parabuthus hamar Kovarík, Lowe, Plísková and Stáhlavsky, 2016 Parabuthus heterurus Pocock, 1899 Parabuthus hunteri Pocock, 1895 Parabuthus kabateki Kovarík, Lowe, Elmi and Stáhlavsky, 2019 Parabuthus kajibu Kovarík, Lowe, Plísková and Stáhlavsky, 2016 Parabuthus kalaharicus Lamoral, 1977 Parabuthus kraepelini Werner, 1902 Parabuthus kuanyamarum (Monard, 1937) Parabuthus laevifrons (Simon, 1888) Parabuthus liosoma (Ehrenberg, 1828) Parabuthus maximus Werner, 1913 Parabuthus mazuchi Kovarík, Lowe, Elmi and Stáhlavsky, 2019 Parabuthus mossambicensis (Peters, 1861) Parabuthus muelleri Prendini, 2000 Parabuthus namibensis Lamoral, 1979 Parabuthus nanus Lamoral, 1979 Parabuthus pallidus Pocock, 1895 Parabuthus planicauda (Pocock, 1889) Parabuthus raudus (Simon, 1888) Parabuthus robustus Kovarík, Lowe, Elmi and Stáhlavsky, 2019 Parabuthus schlechteri Purcell, 1899 Parabuthus setiventer Prendini and Esposito, 2010 Parabuthus somalilandus Kovarík, Lowe, Elmi and Stáhlavsky, 2019 Parabuthus stridulus Hewitt, 1913 Parabuthus transvaalicus Purcell, 1899 Parabuthus villosus (Peters, 1862) Parascorpiops montanus Banks, 1928 Paravaejovis confusus (Stahnke, 1940) Paravaejovis diazi (Williams, 1970) Paravaejovis eusthenura (Wood, 1863) Paravaejovis flavus (Banks, 1900) Paravaejovis galbus (Williams, 1970) Paravaejovis gravicaudus (Williams, 1970) Paravaejovis hoffmanni (Williams, 1970) Paravaejovis pumilis (Williams, 1970) Paravaejovis puritanus (Gertsch, 1958) Paravaejovis schwenkmeyeri (Williams, 1970) Paravaejovis spinigerus (Wood, 1863) Paravaejovis waeringi (Williams, 1970) Paruroctonus ammonastes Haradon, 1984 Paruroctonus arenicola Haradon, 1984 Paruroctonus arnaudi Williams, 1972 Paruroctonus baergi (Williams and Hadley, 1967) Paruroctonus bajae Williams, 1972 Paruroctonus bantai (Gertsch and Soleglad, 1966) Paruroctonus becki (Gertsch and Allred, 1965) Paruroctonus boquillas Sissom and Henson, 1998 Paruroctonus boreus (Girard, 1854) Paruroctonus borregoensis Williams, 1972 Paruroctonus coahuilanus Haradon, 1985 Paruroctonus conclusus Jain, Forbes and Esposito, 2022 Paruroctonus gracilior (Hoffmann, 1931) Paruroctonus hirsutipes Haradon, 1984 Paruroctonus luteolus (Gertsch and Soleglad, 1966) Paruroctonus maritimus Williams, 1987 Paruroctonus marksi Haradon, 1984 Paruroctonus nitidus Haradon, 1984 Paruroctonus pecos Sissom and Francke, 1981 Paruroctonus pseudopumilis (Williams, 1970) Paruroctonus shulovi (Williams, 1970) Paruroctonus silvestrii (Borelli, 1909) Paruroctonus simulatus Haradon, 1985 Paruroctonus soda Jain, Forbes and Esposito, 2022 Paruroctonus stahnkei (Gertsch and Soleglad, 1966) Paruroctonus surensis Williams and Haradon in Williams, 1980 Paruroctonus utahensis (Williams, 1968) Paruroctonus variabilis Hjelle, 1982 Paruroctonus ventosus Williams, 1972 Paruroctonus williamsi Sissom and Francke, 1981 Paruroctonus xanthus (Gertsch and Soleglad, 1966) Pectinibuthus birulai Fet in Orlov and Vasilyev, 1984 Phoniocercus pictus Pocock, 1893 Phoniocercus sanmartini Cekalovic, 1968 Physoctonus amazonicus Lourenço, 2017 Physoctonus debilis (C. L. Koch, 1840) Physoctonus striatus Esposito, Yamaguti, Souza, Pinto-da-Rocha and Prendini, 2017 Picobuthus dundoni Lowe, 2010 Picobuthus wahibaensis Lowe, 2010 Plesiobuthus paradoxus Pocock, 1900 Plesiochactas dilutus (Karsch, 1881) Plesiochactas mitchelli Soleglad, 1976 Plesiochactas vasquezi Trujillo and Armas, 2012 Polisius persicus Fet, Capes and Sissom, 2001 Protoiurus asiaticus (Birula, 1903) Pseudochactas mischi Soleglad, Kovarík and Fet, 2012 Pseudochactas ovchinnikovi Gromov, 1998 Pseudolissothus pusillus Lourenço, 2001 Pseudolychas ochraceus (Hirst, 1911) Pseudolychas pegleri (Purcell, 1901) Pseudolychas transvaalicus Lawrence, 1961 Pseudouroctonus apacheanus (Gertsch and Soleglad, 1972) Pseudouroctonus brysoni Ayrey and Soleglad, 2017 Pseudouroctonus cazieri (Gertsch and Soleglad, 1972) Pseudouroctonus chicano (Gertsch and Soleglad, 1972) Pseudouroctonus kremani Ayrey and Soleglad, 2015 Pseudouroctonus lindsayi (Gertsch and Soleglad, 1972) Pseudouroctonus moyeri Ayrey, Kovarík and Myers, 2021 Pseudouroctonus peccatum Tate, Riddle, Soleglad and Graham, 2013 Pseudouroctonus reddelli (Gertsch and Soleglad, 1972) Pseudouroctonus rufulus (Gertsch and Soleglad, 1972) Pseudouroctonus santarita Ayrey and Soleglad, 2015 Pseudouroctonus sprousei Francke and Savary, 2006 Pseudouroplectes betschi Lourenço, 1995 Pseudouroplectes jacki Lourenço, 2021 Pseudouroplectes lalyae Lourenço and Ythier, 2010 Pseudouroplectes maculatus Lourenço and Goodman, 2006 Pseudouroplectes pidgeoni Lourenço and Goodman, 1999 Pseudouroplectes tsingy Lourenço, Wilmé and Waeber, 2015 Qianxie solegladi Tang, 2022 Razianus birulai Tahir, Navidpour and Prendini, 2014 Razianus farzanpayi Tahir, Navidpour and Prendini, 2014 Razianus xinjianganus Lourenço, Sun and Zhu, 2010 Razianus zarudnyi (Birula, 1903) Reddyanus aareyensis (Mirza and Sanap, 2010) Reddyanus acanthurus (Pocock, 1899) Reddyanus assamensis (Oates, 1888) Reddyanus basilicus (Karsch, 1879) Reddyanus besucheti (Vachon, 1982) Reddyanus bilyi (Kovarík, 2003) Reddyanus brachycentrus (Pocock, 1899) Reddyanus ceylonensis Kovarík, Lowe, Ranawana, Hoferek, Jayarathne, Plísková and Stáhlavsky, 2016 Reddyanus corbeti (Tikader and Bastawade, 1983) Reddyanus deharvengi (Lourenço and Duhem, 2010) Reddyanus feti (Kovarík, 2013) Reddyanus furai Kovarík and Stáhlavsky, 2019 Reddyanus heimi (Vachon, 1976) Reddyanus hofereki Kovarík and Stáhlavsky, 2019 Reddyanus jayarathnei Kovarík in Kovarík et al., 2016 Reddyanus jendeki (Kovarík, 2013) Reddyanus justi Kovarík, Lowe and Stáhlavsky, 2020 Reddyanus khammamensis (Kovarík, 2003) Reddyanus krasenskyi (Kovarík, 1998) Reddyanus kurkai (Kovarík, 1997) Reddyanus loebli (Vachon, 1982) Reddyanus majkusi Kovarík and Stáhlavsky, 2019 Reddyanus melanodactylus (L. Koch, 1867) Reddyanus navaiae (Kovarík, 1998) Reddyanus neradi (Kovarík, 2013) Reddyanus petrzelkai (Kovarík, 2003) Reddyanus problematicus (Kovarík, 2003) Reddyanus ranawanai Kovarík in Kovarík et al., 2016 Reddyanus rigidulus (Pocock, 1897) Reddyanus rolciki Kovarík and Stáhlavsky, 2019 Reddyanus schwotti Kovarík and Stáhlavsky, 2019 Reddyanus tibetanus (Lourenço and Zhu, 2008) Reddyanus vittatus (Pocock, 1900) Reddyanus zideki (Kovarík, 1994) Rhopalurus caribensis Teruel and Roncallo, 2008 Rhopalurus laticauda Thorell, 1876 Rhopalurus ochoai Esposito, Yamaguti, Souza, Pinto-da-Rocha and Prendini, 2017 Rhopalurus renelauerae Lourenço, 2016 Rugodentus keralaensis Bastawade, Sureshan and Radhakrishnan, 2005 Rumikiru atacama Ojanguren-Affilastro, Mattoni, Ochoa and Prendini, 2012 Rumikiru lourencoi (Ojanguren-Affilastro, 2003) Saharobuthus elegans Lourenço and Duhem, 2009 Sahyadrimetrus barberi (Pocock, 1900) Sahyadrimetrus kanarensis (Pocock, 1900) Sahyadrimetrus mathewi Prendini and Loria, 2020 Sahyadrimetrus rugosus (Couzijn, 1981) Sahyadrimetrus scaber (Thorell, 1876) Sahyadrimetrus tikaderi Prendini and Loria, 2020 Sardoscorpius troglophilus Tropea and Onnis, 2020 Sassanidothus gracilis (Birula, 1900) Sassanidothus zarudnyi (Birula, 1900) Scorpio atlasensis Khammassi et al., 2023 Scorpio birulai Fet, 1997 Scorpio ennedi Lourenço, Duhem and Cloudsley-Thompson, 2012 Scorpio fuliginosus (Pallary, 1928) Scorpio fuscus (Ehrenberg, 1829) Scorpio granulomanus Al-Saraireh, Yagmur, Afifeh and Amr, 2023 Scorpio hesperus Birula, 1910 Scorpio iznassen Ythier and Francois, 2023 Scorpio kruglovi Birula, 1910 Scorpio maurus Linnaeus, 1758 Scorpio mogadorensis Birula, 1910 Scorpio moulouya Ythier and Francois, 2023 Scorpio niger Lourenço and Cloudsley-Thompson, 2012 Scorpio occidentalis Werner, 1936 Scorpio palmatus (Ehrenberg, 1828) Scorpio propinquus (Simon, 1872) Scorpio punicus Fet in Fet, Sissom, Lowe and Braunwalder, 2000 Scorpio savanicola Lourenço, 2009 Scorpio schweiggeri Holl, 1829 Scorpio tassili Lourenço and Rossi, 2016 Scorpio touili Ythier and Francois, 2023 Scorpio weidholzi Werner, 1929 Scorpiops afghanus Lourenço and Qi, 2006 Scorpiops alexandreanneorum (Lourenço, 2013) Scorpiops anthracinus Simon, 1887 Scorpiops artemisae (Kovarík, Kosulic, Stáhlavsky, Plísková, Dongkhamfu and Wongprom, 2015) Scorpiops asthenurus Pocock, 1900 Scorpiops atomatus Qi, Zhu and Lourenço, 2005 Scorpiops bastawadei Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops beccaloniae (Kovarík, 2005) Scorpiops bhutanensis Tikader and Bastawade, 1983 Scorpiops binghamii Pocock, 1893 Scorpiops birulai Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops braunwalderi Kovarík, 2000 Scorpiops calmonti (Lourenço, 2013) Scorpiops cavernicola (Lourenço and Pham, 2013) Scorpiops chiangmai (Lourenço, 2019) Scorpiops ciki Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops citadelle (Kovarík, 2013) Scorpiops dakrong (Lourenço and Pham, 2014) Scorpiops dastychi Kovarík, 2000 Scorpiops deccanensis Tikader and Bastawade, 1977 Scorpiops demisi Kovarík, 2005 Scorpiops dii Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops dunlopi Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops farkaci Kovarík, 1993 Scorpiops feti Kovarík, 2000 Scorpiops furai Kovarík, 2020 Scorpiops grandjeani (Vachon, 1974) Scorpiops grosseri Kovarík, 2020 Scorpiops hardwickii (Gervais, 1843) Scorpiops harmsi Kovarík, 2020 Scorpiops hofereki Kovarík, 2020 Scorpiops ingens Yin, Zhang, Pan, Li and Di, 2015 Scorpiops irenae Kovarík, 1994 Scorpiops jendeki Kovarík, 1994 Scorpiops kaftani Kovarík, 1993 Scorpiops kamengensis (Bastawade, 2006) Scorpiops kautti Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops kejvali Kovarík, 2020 Scorpiops krabiensis Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops kubani (Kovarík, 2004) Scorpiops langxian Zhu, Qi and Lourenço in Qi et al., 2005 Scorpiops leptochirus Pocock, 1893 Scorpiops lhasa Di and Zhu, 2009 Scorpiops lii (Di and Qiao, 2020) Scorpiops lindbergi Vachon, 1980 Scorpiops lioneli Sulakhe, Deshpande, Dandekar, Padhye and Bastawade, 2021 Scorpiops longimanus Pocock, 1893 Scorpiops lourencoi Lv and Di, 2022 Scorpiops lowei Tang, 2022 Scorpiops luridus Zhu, Lourenço and Qi in Qi et al., 2005 Scorpiops maharashtraensis (Mirza, Sanap and Upadhye, 2014) Scorpiops margerisonae Kovarík, 2000 Scorpiops montanus Karsch, 1879 Scorpiops nagphani Sulakhe, Deshpande, Dandekar, Padhye and Bastawade, 2021 Scorpiops neera Sulakhe, Deshpande, Dandekar, Padhye and Bastawade, 2021 Scorpiops neradi (Kovarík, Plísková and Stáhlavsky, 2013) Scorpiops novaki (Kovarík, 2005) Scorpiops oligotrichus Fage, 1933 Scorpiops orioni (Kovarík, Kosulic, Stáhlavsky, Plísková, Dongkhamfu and Wongprom, 2015) Scorpiops pachmarhicus Bastawade, 1992 Scorpiops pakistanus Kovarík and Ahmed, 2009 Scorpiops pakseensis Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops petersii Pocock, 1893 Scorpiops phaltanensis (Sulakhe et al., 2020) Scorpiops phatoensis Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops piceus Lourenço and Ythier, 2022 Scorpiops prasiti Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops problematicus Kovarík, 2000 Scorpiops profusus (Lourenço, 2017) Scorpiops pseudomontanus Kovarík and Ahmed, 2009 Scorpiops puerensis (Di, Wu, Cao, Xiao and Li, 2010) Scorpiops rohtangensis Mani, 1959 Scorpiops satarensis Pocock, 1900 Scorpiops scheibeae Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops schumacheri Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops sejnai Kovarík, 2000 Scorpiops sherwoodae Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops shidian (Zhu, Qi and Lourenço in Qi et al., 2005) Scorpiops solegladi Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops solidus Karsch, 1879 Scorpiops songi Di and Qiao, 2020 Scorpiops spitiensis Zambre, Sanap and Mirza, 2014 Scorpiops taxkorgan Lourenço, 2018 Scorpiops telbaila Sulakhe, Deshpande, Dandekar, Ketkar, Padhye and Bastawade, 2020 Scorpiops tenuicauda Pocock, 1894 Scorpiops thailandus Kovarík, Lowe, Stockmann and Stáhlavsky, 2020 Scorpiops thaomischi (Kovarík, 2012) Scorpiops tibetanus Hirst, 1911 Scorpiops tongtongi Tang, 2022 Scorpiops troglodytes (Lourenço and Pham, 2015) Scorpiops tryznai Kovarík, 2020 Scorpiops vachoni (Zhu, Qi and Lourenço in Qi et al., 2005) Scorpiops validus (Di et al., 2010) Scorpiops viktoriae (Lourenço and Kosulic, 2018) Scorpiops vonwicki Birula, 1913 Scorpiops vrushchik Sulakhe, Deshpande, Dandekar, Padhye and Bastawade, 2021 Scorpiops wongpromi (Kovarík, Soleglad and Kosulic, 2013) Scorpiops wrzecionkoi Kovarík, 2020 Scorpiops xui (Sun and Zhu, 2010) Scorpiops yagmuri Kovarík, 2020 Scorpiops yangi (Zhu, Zhang and Lourenço, 2007) Scorpiops zhangshuyuani (Ythier, 2019) Scorpiops zhui Lv, Lourenço and Di, 2023 Scorpiops zubairahmedi Kovarík in Kovarík and Ahmed, 2009 Scorpiops zubairi Kovarík, 2020 Serradigitus adcocki (Williams, 1980) Serradigitus armadentis (Williams, 1980) Serradigitus baueri (Gertsch, 1958) Serradigitus bechteli (Williams, 1980) Serradigitus calidus (Soleglad, 1974) Serradigitus dwyeri (Williams, 1980) Serradigitus gertschi (Williams, 1968) Serradigitus gigantaensis (Williams, 1980) Serradigitus gramenestris (Williams, 1970) Serradigitus haradoni (Williams, 1980) Serradigitus hearnei (Williams, 1980) Serradigitus joshuaensis (Soleglad, 1972) Serradigitus littoralis (Williams, 1980) Serradigitus minutus (Williams, 1970) Serradigitus miscionei Ayrey, 2011 Serradigitus pacificus (Williams, 1980) Serradigitus torridus Williams and Berke, 1986 Serradigitus wupatkiensis (Stahnke, 1940) Serradigitus yaqui Sissom and Stockwell, 1991 Sinoscorpius shandongensis Hong, 1983 Smeringurus aridus (Soleglad, 1972) Smeringurus grandis (Williams, 1970) Smeringurus mesaensis (Stahnke, 1957) Smeringurus vachoni (Stahnke, 1961) Somalibuthus demisi Kovarík, 1998 Somalibuthus sabae Kovarík and Njoroge, 2021 Somalicharmus whitmanae Kovarík, 1998 Sotanochactas elliotti (Mitchell, 1971) Spelaeolychas hosei (Pocock, 1891) Spinoburmesebuthus knodelorum Lourenço, 2021 Spinoburmesebuthus pohli Lourenço in Lourenço and Velten, 2017 Spinoburmesebuthus spinipedis (Xuan, Cai and Huang, 2022) Spinochactas camopi Lourenço, Chevalier and Ythier, 2022 Spinochactas mitaraka Lourenço, 2016 Srilankametrus caesar (C. L. Koch, 1841) Srilankametrus couzijni Prendini and Loria, 2020 Srilankametrus gravimanus (Pocock, 1894) Srilankametrus indus (De Geer, 1778) Srilankametrus pococki Prendini and Loria, 2020 Srilankametrus serratus (Pocock, 1900) Srilankametrus yaleensis (Kovarík et al., 2019) Stahnkeus allredi (Sissom and Stockwell, 1991) Stahnkeus deserticola (Williams, 1970) Stahnkeus harbisoni (Williams, 1970) Stahnkeus polisi (Sissom and Stockwell, 1991) Stahnkeus subtilimanus (Soleglad, 1972) Stygochactas granulosus (Sissom and Cokendolpher, 1998) Sucinlourencous adrianae Rossi, 2015 Superstitionia donensis Stahnke, 1940 Syntropis aalbui Lowe, Soleglad and Fet in Soleglad et al., 2007 Syntropis macrura Kraepelin, 1900 Syntropis williamsi Soleglad, Lowe and Fet, 2007 Tarsoporosus anchicaya Lourenço and Flórez, 1990 Tarsoporosus flavus (González-Sponga, 1984) Tarsoporosus kugleri (Schenkel, 1932) Tarsoporosus macuira Teruel and Roncallo, 2007 Tarsoporosus tizopedrosoi Bedoya-Roqueme and Lira, 2022 Tarsoporosus yustizi (González-Sponga, 1984) Tehuankea moyanoi Cekalovic, 1973 Teruelius ankarafantsika (Lourenço, 2003) Teruelius ankarana (Lourenço and Goodman, 2003) Teruelius annulatus (Fage, 1929) Teruelius bemaraha (Lourenço, Wilmé and Waeber, 2018) Teruelius bicolor (Lourenço, 2012) Teruelius bistriatus (Kraepelin, 1900) Teruelius eliseanneae (Lourenço and Wilmé, 2016) Teruelius feti (Lourenço, 1996) Teruelius flavopiceus (Kraepelin, 1900) Teruelius ganzhorni (Lourenço, Wilmé and Waeber, 2016) Teruelius grandidieri (Kraepelin, 1900) Teruelius haeckeli Lowe and Kovarík, 2022 Teruelius intertidalis (Lourenço, 1999) Teruelius limbatus (Pocock, 1889) Teruelius magalieae (Lourenço, 2014) Teruelius mahafaliensis (Lourenço, Goodman and Ramilijaona, 2004) Teruelius makay (Lourenço and Wilmé, 2015) Teruelius mavo (Lourenço and Rossi in Lourenço et al., 2020) Teruelius olgae (Lourenço, 2004) Teruelius rossii (Lourenço, 2013) Teruelius sabineae (Lourenço and Wilmé, 2016) Teruelius waeberi (Lourenço and Wilmé, 2016) Tetratrichobothrius flavicaudis (De Geer, 1778) Teuthraustes adrianae González-Sponga, 1975 Teuthraustes akananensis González-Sponga, 1984 Teuthraustes amazonicus (Simon, 1880) Teuthraustes atramentarius Simon, 1878 Teuthraustes braziliensis Lourenço and Duhem, 2010 Teuthraustes camposi (Mello-Leitão, 1939) Teuthraustes carmelinae Scorza, 1954 Teuthraustes castiglii Rossi, 2015 Teuthraustes dubius (Borelli, 1899) Teuthraustes festae (Borelli, 1899) Teuthraustes gervaisii (Pocock, 1893) Teuthraustes giupponii Ythier and Lourenço, 2017 Teuthraustes glaber Kraepelin, 1912 Teuthraustes guerdouxi Lourenço, 1995 Teuthraustes japura Lourenço and Ythier, 2022 Teuthraustes khodayarii Ythier and Lourenço, 2017 Teuthraustes kuryi Ythier and Lourenço, 2017 Teuthraustes lisei Lourenço, 1994 Teuthraustes lojanus (Pocock, 1900) Teuthraustes maturaca González-Sponga, 1991 Teuthraustes newaribe Lourenço, Giupponi and Pedroso, 2011 Teuthraustes oculatus Pocock, 1900 Teuthraustes ohausi Kraepelin, 1912 Teuthraustes reticulatus González-Sponga, 1991 Teuthraustes rosenbergi (Pocock, 1898) Teuthraustes simonsi (Pocock, 1900) Teuthraustes whymperi (Pocock, 1893) Teuthraustes wittii (Kraepelin, 1896) Thaicharmus guptai Mirza, Sanap and Kunte, 2016 Thaicharmus indicus Kovarík, 2013 Thaicharmus lowei Kovarík, Soleglad and Fet, 2007 Thaicharmus mahunkai Kovarík, 1995 Thestylus aurantiurus Yamaguti and Pinto-da-Rocha, 2003 Thestylus glasioui (Bertkau, 1880) Thestylus signatus Mello-Leitão, 1931 Thorellius cristimanus (Pocock, 1898) Thorellius intrepidus (Thorell, 1876) Thorellius tekuani González-Santillán and Prendini, 2018 Thorellius wixarika González-Santillán and Prendini, 2018 Thorellius yuyuawi González-Santillán and Prendini, 2018 Tibetiomachus himalayensis Lourenço and Qi, 2006 Timogenes dorbignyi (Guérin-Méneville, 1843) Timogenes elegans (Mello-Leitão, 1931) Timogenes haplochirus Maury and Roig Alsina, 1977 Timogenes mapuche Maury, 1975 Timogenes sumatranus Simon, 1880 Tityobuthus antsingy Lourenço and Goodman, 2004 Tityobuthus baroni (Pocock, 1890) Tityobuthus betschi Lourenço, Qi and Goodman, 2008 Tityobuthus chelbergorum Lourenço, Qi and Goodman, 2008 Tityobuthus darainensis Lourenço and Goodman, 2002 Tityobuthus dastychi Lourenço, 1997 Tityobuthus griswoldi Lourenço, 2000 Tityobuthus guillaumeti Lourenço, 1995 Tityobuthus ivohibe Lourenço and Goodman, 1999 Tityobuthus judsoni Lourenço, 1996 Tityobuthus lokobe Lourenço, Waeber and Wilmé, 2016 Tityobuthus maccarteri Lourenço, Qi and Goodman, 2008 Tityobuthus manonae Lourenço, 2000 Tityobuthus mariejeanneae Lourenço, Waeber and Wilmé, 2018 Tityobuthus monodi Lourenço, 2000 Tityobuthus orangea Lourenço, Waeber and Wilmé, 2020 Tityobuthus pallidus Lourenço, 2004 Tityobuthus parrilloi Lourenço, 1996 Tityobuthus petrae Lourenço, 1996 Tityobuthus pococki Lourenço, 1995 Tityobuthus rakotondravonyi Lourenço and Goodman, 2003 Tityopsis canizaresorum Teruel and Rodríguez-Cabrera, 2020 Tityopsis inaequalis (Armas, 1974) Tityopsis inexpectata (Moreno, 1940) Tityopsis mulata Teruel and Rodríguez-Cabrera, 2020 Tityopsis pumila Teruel and Rodríguez-Cabrera, 2020 Tityopsis sheylae Teruel and Rodríguez-Cabrera, 2020 Tityus aba Candido, Lucas, Souza, Dias and Lira-da-Silva, 2005 Tityus abudi Armas, 1999 Tityus acananensis González-Sponga, 2009 Tityus adisi Lourenço and Pézier, 2002 Tityus adrianoi Lourenço, 2003 Tityus ahincoi González-Sponga, 2001 Tityus alejandroi Teruel, Rivera and Santos, 2015 Tityus altithronus Armas, 1999 Tityus androcottoides (Karsch, 1879) Tityus anduzei González-Sponga, 1997 Tityus angelesae Santiago-Blay, 2009 Tityus anneae Lourenço, 1997 Tityus anori Lourenço, Rossi and Wilmé, 2019 Tityus antioquensis Lourenço and Otero Patiño, 1998 Tityus apiacas Lourenço, 2002 Tityus apozonalli Riquelme, Villegas and González in Riquelme et al., 2015 Tityus arellanoparrai González-Sponga, 1985 Tityus argentinus Borelli, 1899 Tityus asthenes Pocock, 1893 Tityus atriventer Pocock, 1897 Tityus bahiensis (Perty, 1833) Tityus barquisimetanus González-Sponga, 1994 Tityus bastosi Lourenço, 1984 Tityus bellulus Armas, 1999 Tityus birabeni Abalos, 1955 Tityus blanci Lourenço, 1994 Tityus blaseri Mello-Leitão, 1931 Tityus boconoensis González-Sponga, 1981 Tityus bolivianus Kraepelin, 1895 Tityus brazilae Lourenço and Eickstedt, 1984 Tityus breweri González-Sponga, 1997 Tityus cachipalensis González-Sponga, 2002 Tityus caesarbarrioi González-Sponga, 2001 Tityus canopensis Lourenço and Pézier, 2002 Tityus carabobensis González-Sponga, 1987 Tityus carinatoides Mello-Leitão, 1945 Tityus caripitensis Quiroga, de Sousa and Parrilla-Alvarez, 2000 Tityus carolinae Kovarík, Teruel, Cozijn and Seiter, 2013 Tityus carrilloi Ojanguren-Affilastro in Ojanguren-Affilastro et al., 2021 Tityus carvalhoi Mello-Leitão, 1945 Tityus cerroazul Lourenço, 1986 Tityus championi Pocock, 1898 Tityus charalaensis Mello-Leitão, 1940 Tityus charreyroni Vellard, 1932 Tityus chilensis Lourenço, 2005 Tityus choco Lourenço and Flórez, 2018 Tityus cisandinus Lourenço and Ythier, 2017 Tityus clathratus C. L. Koch, 1844 Tityus columbianus (Thorell, 1876) Tityus confluens Borelli, 1899 Tityus costatus (Karsch, 1879) Tityus crassicauda Lourenço and Ythier, 2013 Tityus crassimanus (Thorell, 1876) Tityus cuellari Lourenço, 1994 Tityus culebrensis González-Sponga, 1994 Tityus curupi Ojanguren-Affilastro, Adilardi, Cajade, Ramírez, Ceccarelli and Mola, 2017 Tityus cylindricus (Karsch, 1879) Tityus dasyurus Pocock, 1897 Tityus dedoslargos Francke and Stockwell, 1987 Tityus demangei Lourenço, 1981 Tityus dinizi Lourenço, 1997 Tityus discrepans (Karsch, 1879) Tityus dorae González-Sponga, 2001 Tityus duacaensis González-Sponga, 2007 Tityus dulceae González-Sponga, 2006 Tityus dupouyi González-Sponga, 1987 Tityus ecuadorensis Kraepelin, 1896 Tityus elii Armas and Marcano Fondeur, 1992 Tityus elizabethae Lourenço and Ramos, 2004 Tityus elizabethebravoi González-Sponga and Wall González, 2007 Tityus engelkei Pocock, 1902 Tityus eogenus Menge, 1869 Tityus estherae Santiago-Blay, 2009 Tityus evandroi Mello-Leitão, 1945 Tityus exstinctus Lourenço, 1995 Tityus fasciolatus Pessôa, 1935 Tityus festae Borelli, 1899 Tityus filodendron González-Sponga, 1981 Tityus florezi Lourenço, 2000 Tityus footei Chamberlin, 1916 Tityus forcipula (Gervais, 1843) Tityus fuhrmanni Kraepelin, 1914 Tityus funestus Hirst, 1911 Tityus gaffini Lourenço, 2000 Tityus gasci Lourenço, 1982 Tityus generaltheophiloi Lourenço, 2017 Tityus geratus Santiago-Blay and Poinar, 1988 Tityus gonzalespongai Quiroga, de Sousa, Parrilla-Álvarez and Manzanilla, 2004 Tityus grahami Lourenço, 2012 Tityus grottoedensis Botero-Trujillo and Flórez, 2014 Tityus guane Moreno-González, González and Flórez, 2019 Tityus guaricoensis González-Sponga, 2004 Tityus haetianus Teruel and Santos, 2018 Tityus horacioi Lourenço and Leguin, 2011 Tityus imei Borges, de Sousa and Manzanilla, 2006 Tityus indecisus Mello-Leitão, 1934 Tityus insignis (Pocock, 1889) Tityus intermedius Borelli, 1899 Tityus isabelceciliae González-Sponga, D'Suze and Sevcik, 2001 Tityus ivani González-Sponga, 2008 Tityus ivicnancor González-Sponga, 1997 Tityus jaimei Miranda, Bermúdez, Flórez D. and Armas, 2020 Tityus jeanvellardi Lourenço, 2001 Tityus julianae Lourenço, 2005 Tityus juliorum Santiago-Blay, 2009 Tityus jussarae Lourenço, 1988 Tityus kaderkai Kovarík, 2005 Tityus kalettai González-Sponga, 2007 Tityus karaja Lourenço, 2016 Tityus kindli Kovarík and Teruel, 2014 Tityus knodeli Lourenço, 2014 Tityus kukututee Ythier, Chevalier and Gangadin, 2020 Tityus kuryi Lourenço, 1997 Tityus lancinii González-Sponga, 1972 Tityus lokiae Lourenço, Adis and Araujo, 2005 Tityus longidigitus González-Sponga, 2008 Tityus lourencoi Flórez, 1996 Tityus lutzi Giltay, 1928 Tityus macrochirus Pocock, 1897 Tityus magnimanus Pocock, 1897 Tityus maimirensis González-Sponga, 2007 Tityus mana Lourenço, 2012 Tityus manakai González-Sponga, 2004 Tityus maniapurensis González-Sponga, 2009 Tityus marajoensis Lourenço and da Silva, 2007 Tityus maranhensis Lourenço, Junior and Limeira-de-Oliveira, 2006 Tityus marechali Lourenço, 2013 Tityus martinpaechi Lourenço, 2001 Tityus matthieseni Pinto-da-Rocha and Lourenço, 2000 Tityus mattogrossensis Borelli, 1901 Tityus maturinensis González-Sponga, 2008 Tityus melanostictus Pocock, 1893 Tityus melici Lourenço, 2003 Tityus metuendus Pocock, 1897 Tityus michelii Armas, 1982 Tityus microcystis Lutz and Mello, 1922 Tityus monaguensis González-Sponga, 1974 Tityus mongei Lourenço, 1996 Tityus moralensis Moreno-González, Pinto-da-Rocha and Cabra-García, 2022 Tityus mraceki Kovarík, 2007 Tityus mucusunamensis González-Sponga, 2006 Tityus munozi Lourenço, 1997 Tityus neblina Lourenço, 2008 Tityus neglectus Mello-Leitão, 1932 Tityus neibae Armas, 1999 Tityus nelsoni Lourenço, 2005 Tityus nematochirus Mello-Leitão, 1940 Tityus neoespartanus González-Sponga, 1996 Tityus nororientalis González-Sponga, 1996 Tityus obispoi González-Sponga, 2006 Tityus obscurus (Gervais, 1843) Tityus obtusus (Karsch, 1879) Tityus ocelote Francke and Stockwell, 1987 Tityus osmanus González-Sponga, 1996 Tityus oteroi Lourenço, 1998 Tityus ottenwalderi Armas, 1999 Tityus pachyurus Pocock, 1897 Tityus pampanensis González-Sponga, 2007 Tityus panguana Kovarík, Teruel, Lowe and Friedrich, 2015 Tityus paraguayensis Kraepelin, 1895 Tityus parvulus Kraepelin, 1914 Tityus paulistorum Lourenço and da Silva, 2006 Tityus perijanensis González-Sponga, 1994 Tityus pictus Pocock, 1893 Tityus pintodarochai Lourenço, 2005 Tityus pittieri González-Sponga, 1981 Tityus pococki Hirst, 1907 Tityus portoplatensis Armas and Marcano Fondeur, 1992 Tityus potameis Lourenço and Giupponi, 2004 Tityus prancei Lourenço, 2000 Tityus proseni Abalos, 1954 Tityus pugilator Pocock, 1898 Tityus pusillus Pocock, 1893 Tityus quiriquirensis González-Sponga, 2008 Tityus quirogae De Sousa, Manzanilla and Parrilla-Álvarez, 2006 Tityus quisqueyanus Armas, 1982 Tityus ramirezi Esquivel de Verde, 1969 Tityus raquelae Lourenço, 1988 Tityus rebieri Lourenço, 1997 Tityus riocaurensis González-Sponga, 1996 Tityus rionegrensis Lourenço, 2006 Tityus riverai Teruel and Sánchez, 2009 Tityus roigi Maury and Lourenço, 1987 Tityus rojasi González-Sponga, 1996 Tityus romeroi González-Sponga, 2008 Tityus rondonorum Rojas-Runjaic and Armas, 2007 Tityus rosenbergi Pocock, 1898 Tityus rufofuscus Pocock, 1897 Tityus rugosus Schenkel, 1932 Tityus rupestre Lourenço, 2019 Tityus rusmelyae González-Sponga, D'Suze and Sevcik, 2001 Tityus sabineae Lourenço, 1994 Tityus sanarensis González-Sponga, 1997 Tityus sanfernandoi González-Sponga, 2008 Tityus sarisarinamensis González-Sponga, 2002 Tityus sastrei Lourenço and Flórez, 1989 Tityus schrammi Teruel and Santos, 2018 Tityus serrulatus Lutz and Mello, 1922 Tityus shiriana González-Sponga, 1991 Tityus silvestris Pocock, 1897 Tityus simonsi Pocock, 1900 Tityus smithii Pocock, 1893 Tityus soratensis Kraepelin, 1912 Tityus spelaeus Moreno-González, Pinto-da-Rocha and Gallão, 2021 Tityus stigmurus (Thorell, 1876) Tityus strandi Werner, 1939 Tityus surmeridensis González-Sponga, 2002 Tityus surorientalis González-Sponga, 1996 Tityus sylviae Lourenço, 2005 Tityus tamayoi González-Sponga, 1987 Tityus tayrona Lourenço, 1991 Tityus tenuicauda Prendini, 2001 Tityus timendus Pocock, 1898 Tityus tocantim Lourenço, 2022 Tityus trinitatis Pocock, 1897 Tityus trivittatus Kraepelin, 1898 Tityus tucurui Lourenço, 1988 Tityus uniformis Mello-Leitão, 1931 Tityus unus Pinto-da-Rocha and Lourenço, 2000 Tityus uquirensis González-Sponga, 2001 Tityus urachichensis González-Sponga, 2007 Tityus urbinai Scorza, 1952 Tityus uruguayensis Borelli, 1901 Tityus vaissadei Lourenço, 2002 Tityus valerae Scorza, 1954 Tityus venamensis González-Sponga, 1981 Tityus ventuarensis González-Sponga, 2009 Tityus walli González-Sponga and Wall-González, 2007 Tityus yerenai González-Sponga, 2009 Tityus ythieri Lourenço, 2007 Tityus zulianus González-Sponga, 1981 Troglocormus ciego Francke, 1981 Troglocormus willis Francke, 1981 Troglokhammouanus steineri Lourenço, 2007 Troglopolyphemos anophthalmus (Francke, 1977) Troglorhopalurus lacrau (Lourenço and Pinto-da-Rocha, 1997) Troglorhopalurus translucidus Lourenço, Baptista and Giupponi, 2004 Troglotayosicus ballvei Botero-Trujillo, Ochoa and Prendini, 2021 Troglotayosicus hirsutus Botero-Trujillo, Ochoa, Tovar and Souza, 2012 Troglotayosicus humiculum Botero-Trujillo and Francke, 2009 Troglotayosicus meijdeni Botero-Trujillo et al., 2017 Troglotayosicus muranunkae Lourenço, Sánchez-Vialas, Blasco-Aróstegui and García-Gila, 2020 Troglotayosicus vachoni Lourenço, 1981 Troglotityobuthus gracilis (Fage, 1946) Trypanothacus azraqensis Al-Saraireh, Afifeh, Aloufi, Amr and Lourenço, 2021 Trypanothacus barnesi Lowe, Kovarík, Stockmann and Stáhlavsky, 2019 Trypanothacus buettikeri (Hendrixson, 2006) Typhlochactas cavicola Francke, 1986 Typhlochactas mitchelli Sissom, 1988 Typhlochactas reddelli Mitchell, 1968 Typhlochactas rhodesi Mitchell, 1968 Typhlochactas sissomi Francke, Vignoli and Prendini, 2009 Typhlochactas sylvestris Mitchell and Peck, 1977 Uroctonites giulianii Williams and Savary, 1991 Uroctonites huachuca (Gertsch and Soleglad, 1972) Uroctonites montereus (Gertsch and Soleglad, 1972) Uroctonites sequoia (Gertsch and Soleglad, 1972) Uroctonus franckei Williams, 1986 Uroctonus grahami Gertsch and Soleglad, 1972 Uroctonus mordax Thorell, 1876 Urodacus armatus Pocock, 1888 Urodacus butleri Volschenk, Harvey and Prendini, 2012 Urodacus carinatus Hirst, 1911 Urodacus centralis L. E. Koch, 1977 Urodacus elongatus L. E. Koch, 1977 Urodacus excellens Pocock, 1888 Urodacus giulianii L. E. Koch, 1977 Urodacus hartmeyeri Kraepelin, 1908 Urodacus hoplurus Pocock, 1898 Urodacus koolanensis L. E. Koch, 1977 Urodacus lowei L. E. Koch, 1977 Urodacus macrurus Pocock, 1899 Urodacus manicatus (Thorell, 1876) Urodacus mckenziei Volschenk, Smith and Harvey, 2000 Urodacus megamastigus L. E. Koch, 1977 Urodacus novaehollandiae Peters, 1861 Urodacus planimanus Pocock, 1893 Urodacus similis L. E. Koch, 1977 Urodacus spinatus Pocock, 1902 Urodacus varians Glauert, 1963 Urodacus yaschenkoi (Birula, 1903) Urophonius achalensis Abalos and Hominal, 1974 Urophonius araucano Ojanguren-Affilastro in Ojanguren-Affilastro et al., 2020 Urophonius brachycentrus (Thorell, 1876) Urophonius eugenicus (Mello-Leitão, 1931) Urophonius exochus (Penther, 1913) Urophonius granulatus Pocock, 1898 Urophonius iheringi Pocock, 1893 Urophonius mahuidensis Maury, 1973 Urophonius martinezi Ojanguren-Affilastro and Cheli, 2009 Urophonius mondacai Ojanguren-Affilastro, Pizarro-Araya and Prendini, 2011 Urophonius pehuenche Ojanguren-Affilastro and Pizarro-Araya in Ojanguren-Affilastro et al., 2020 Urophonius pizarroi Ojanguren-Affilastro, Ochoa, Mattoni and Prendini, 2010 Urophonius somuncura Acosta, 2003 Urophonius transandinus Acosta, 1998 Urophonius tregualemuensis Cekalovic, 1981 Urophonius tumbensis Cekalovic, 1981 Uroplectes ansiedippenaarae Prendini, 2015 Uroplectes carinatus (Pocock, 1890) Uroplectes chubbi Hirst, 1911 Uroplectes emiliae (Werner, 1916) Uroplectes fischeri (Karsch, 1879) Uroplectes flavoviridis Peters, 1861 Uroplectes formosus Pocock, 1890 Uroplectes gracilior Hewitt, 1914 Uroplectes insignis Pocock, 1890 Uroplectes katangensis Prendini, 2015 Uroplectes lineatus (C. L. Koch, 1844) Uroplectes longimanus Werner, 1936 Uroplectes machadoi Lourenço, 2000 Uroplectes malawicus Prendini, 2015 Uroplectes marlothi Purcell, 1901 Uroplectes ngangelarum Monard, 1930 Uroplectes occidentalis Simon, 1876 Uroplectes olivaceus Pocock, 1896 Uroplectes otjimbinguensis (Karsch, 1879) Uroplectes pardalis Werner, 1913 Uroplectes pardii Kovarík, 2003 Uroplectes pictus Werner, 1913 Uroplectes pilosus (Thorell, 1876) Uroplectes planimanus (Karsch, 1879) Uroplectes schlechteri Purcell, 1901 Uroplectes schubotzi Kraepelin, 1929 Uroplectes silvestrii Borelli, 1913 Uroplectes teretipes Lawrence, 1966 Uroplectes triangulifer (Thorell, 1876) Uroplectes tumidimanus Lamoral, 1979 Uroplectes variegatus (C. L. Koch, 1844) Uroplectes vittatus (Thorell, 1876) Uroplectes zambezicus Prendini, 2015 Vachonia martinezi Abalos, 1954 Vachoniochactas amazonicus González-Sponga, 1991 Vachoniochactas ashleeae Lourenço, 1994 Vachoniochactas humboldti Flórez, Botero-Trujillo and Acosta, 2008 Vachoniochactas lasallei (González-Sponga, 1978) Vachoniochactas roraima Lourenço and Duhem, 2009 Vachoniolus batinahensis Lowe, 2010 Vachoniolus gallagheri Lowe, 2010 Vachoniolus globimanus Levy, Amitai and Shulov, 1973 Vachoniolus iranus Navidpour, Kovarík, Soleglad and Fet, 2008 Vachonus asiyaae Amir and Kamaluddin, 2009 Vachonus inexpectatus Lourenço, 2015 Vachonus iqbali Amir and Kamaluddin, 2009 Vachonus rajasthanicus Tikader and Bastawade, 1983 Vaejovis aguazarca Díaz-Plascencia and González-Santillán in Chávez-Samoyoa et al., 2022 Vaejovis aquascalentensis Chávez-Samayoa and González-Santillán in Chávez-Samayoa et al., 2022 Vaejovis baggins Azzinnari, Bryson, Graham, Solís-Rojas and Sissom, 2021 Vaejovis bandido Graham, Ayrey and Bryson, 2012 Vaejovis bigelowi Sissom, 2011 Vaejovis carolinianus (Palisot de Beauvois, 1805) Vaejovis cashi Graham, 2007 Vaejovis ceboruco Contreras-Félix and Francke, 2019 Vaejovis chiapas Sissom, 1989 Vaejovis chisos Sissom, 1990 Vaejovis coalcoman Contreras-Félix and Francke, 2014 Vaejovis crumpi Ayrey and Soleglad, 2011 Vaejovis curvidigitus Sissom, 1991 Vaejovis darwini Santibañez-López and Francke, 2010 Vaejovis davidi Soleglad and Fet, 2005 Vaejovis deboerae Ayrey, 2009 Vaejovis decipiens Hoffmann, 1931 Vaejovis dugesi Pocock, 1902 Vaejovis dzahui Santibañez-López and Francke, 2010 Vaejovis electrum Hughes, 2011 Vaejovis elii Ayrey, 2020 Vaejovis feti Graham, 2007 Vaejovis franckei Sissom, 1989 Vaejovis gracilis Gertsch and Soleglad, 1972 Vaejovis grahami Ayrey and Soleglad, 2014 Vaejovis granulatus Pocock, 1898 Vaejovis grayae Ayrey, 2014 Vaejovis halli Ayrey, 2012 Vaejovis intermedius Borelli, 1915 Vaejovis islaserrano Barrales-Alcalá, Francke, Devender and Contreras-Félix, 2018 Vaejovis janssi Williams, 1980 Vaejovis jonesi Stahnke, 1940 Vaejovis lapidicola Stahnke, 1940 Vaejovis maculosus Sissom, 1989 Vaejovis mcwesti Sissom, Graham, Donaldson and Bryson Jr., 2016 Vaejovis mendozai Contreras-Félix and Francke, 2021 Vaejovis mexicanus C. L. Koch, 1836 Vaejovis miscionei Myers and Ayrey, 2021 Vaejovis mitchelli Sissom, 1991 Vaejovis montanus Graham and Bryson, 2010 Vaejovis monticola Sissom, 1989 Vaejovis morelia Miranda-López, Ponce-Saavedra and Francke, 2012 Vaejovis nanchititla Contreras-Félix and Francke, 2019 Vaejovis nigrescens Pocock, 1898 Vaejovis nigrofemoratus Hendrixson and Sissom, 2001 Vaejovis norteno Sissom et al., 2004 Vaejovis ocotensis Zárate-Gálvez and Francke, 2009 Vaejovis patagonia Ayrey, 2018 Vaejovis paysonensis Soleglad, 1973 Vaejovis pequeno Hendrixson, 2001 Vaejovis pococki Sissom, 1991 Vaejovis prendinii Santibañez-López and Francke, 2010 Vaejovis pusillus Pocock, 1898 Vaejovis rossmani Sissom, 1989 Vaejovis santibagnezi Contreras-Félix and Francke, 2019 Vaejovis setosus Sissom, 1989 Vaejovis sierrae Sissom, Graham, Donaldson and Bryson Jr., 2016 Vaejovis smithi Pocock, 1902 Vaejovis solegladi Sissom, 1991 Vaejovis sprousei Sissom, 1990 Vaejovis stetsoni Ayrey and Myers, 2019 Vaejovis talpa Contreras-Félix and Francke, 2019 Vaejovis tapalpa Contreras-Félix and Francke, 2019 Vaejovis tenamaztlei Contreras-Félix, Francke and Bryson Jr., 2015 Vaejovis tenuipalpus Sissom, Hughes, Bryson and Prendini, 2012 Vaejovis tesselatus Hendrixson and Sissom, 2001 Vaejovis tilae Contreras-Félix, del-Pozo and Navarrete-Heredia, 2023 Vaejovis trespicos Zárate-Gálvez and Francke, 2009 Vaejovis trinityae Ayrey, 2013 Vaejovis troupi Ayrey and Soleglad, 2015 Vaejovis vaquero Gertsch and Soleglad, 1972 Vaejovis vorhiesi Stahnke, 1940 Vaejovis zapoteca Santibañez-López and Francke, 2010 Vejovoidus longiunguis (Williams, 1969) Vietbocap canhi Lourenço and Pham, 2010 Vizcaino viscainensis (Williams, 1970) Wernerius inyoensis Webber, Graham and Jaeger, 2012 Wernerius mumai (Sissom, 1993) Wernerius spicatus (Haradon, 1974) Xenobuthus anthracinus (Pocock, 1895) Xenobuthus arabicus (Lourenço and Qi, 2006) Xenobuthus xanthus Lowe, 2018 Zabius birabeni Mello-Leitão, 1938 Zabius fuscus (Thorell, 1876) Zabius gaucho Acosta, Candido, Buckup and Brescovit, 2008
Androctonus amoreuxi amoreuxi (Audouin, 1826) Androctonus amoreuxi levyi Fet, 1997 Androctonus bicolor bicolor Ehrenberg, 1828 Androctonus bicolor longecarinatus (Caporiacco, 1932) Anuroctonus pococki bajae Soleglad and Fet, 2004 Anuroctonus pococki pococki Soleglad and Fet, 2004 Babycurus buettneri buettneri Karsch, 1886 Babycurus buettneri savanicola Lourenço, Ramos and Cloudsley-Thompson, 2005 Bothriurus illudens araponguensis Bücherl et al., 1963 Bothriurus illudens illudens Mello-Leitão, 1947 Bothriurus rochai occidentalis Lourenço, 2000 Bothriurus rochai rochai Mello-Leitão, 1932 Buthacus leptochelys granosus Borelli, 1929 Buthacus leptochelys leptochelys (Ehrenberg, 1829) Centruroides elegans edentulus Werner, 1939 Centruroides elegans elegans (Thorell, 1876) Centruroides elegans insularis Pocock, 1902 Centruroides flavopictus flavopictus (Pocock, 1898) Centruroides flavopictus meridionalis Hoffmann, 1932 Centruroides margaritatus margaritatus (Gervais, 1841) Centruroides nitidus nitidus (Thorell, 1876) Centruroides nitidus taino Armas and Marcano Fondeur, 1987 Centruroides suffusus chiaravigli Borelli, 1915 Centruroides suffusus suffusus Pocock, 1902 Cheloctonus anthracinus anthracinus Pocock, 1899 Cheloctonus anthracinus warreni Hewitt, 1931 Cheloctonus crassimanus crassimanus (Pocock, 1896) Cheloctonus crassimanus depressus Hewitt, 1918 Cheloctonus jonesii jonesii Pocock, 1892 Cheloctonus jonesii sculpturatus Hewitt, 1914 Diplocentrus mexicanus karschi Sissom and Francke, 1998 Diplocentrus mexicanus mexicanus Peters, 1861 Euscorpius carpathicus carpathicus (Linnaeus, 1767) Euscorpius carpathicus fanzagoi Simon, 1879 Euscorpius garganicus garganicus Di Caporiacco, 1950 Euscorpius garganicus molisanus Tropea, 2017 Hadogenes trichiurus caffer Hewitt, 1918 Hadogenes trichiurus gracilioides Hewitt, 1918 Hadogenes trichiurus pallidus Pocock, 1898 Hadogenes trichiurus parvus Hewitt, 1925 Hadogenes trichiurus trichiurus (Gervais, 1843) Hadogenes trichiurus werneri Fet, 1997 Hadogenes trichiurus whitei Purcell, 1899 Hadogenes troglodytes crassicaudatus Hewitt, 1918 Hadogenes troglodytes dentatus Hewitt, 1918 Hadogenes troglodytes letabensis Werner, 1933 Hadogenes troglodytes matoppoanus Hewitt, 1918 Hadogenes troglodytes troglodytes (Peters, 1861) Hadrurus arizonensis arizonensis Ewing, 1928 Hadrurus arizonensis austrinus Williams, 1970 Heterometrus liophysa liophysa (Thorell, 1888) Heterometrus longimanus angustimanus (Thorell, 1876) Heterometrus longimanus borneensis (Thorell, 1876) Heterometrus longimanus humilis (Simon, 1877) Heterometrus longimanus longimanus (Herbst, 1800) Heterometrus petersii petersii (Thorell, 1876) Heteronebo jamaicae jamaicae Francke, 1978 Heteronebo jamaicae occidentalis Francke, 1978 Heteronebo jamaicae portlandensis Francke, 1978 Hottentotta alticola alticola (Pocock, 1895) Hottentotta alticola nigrifrons (Pocock, 1900) Hottentotta minax minax (L. Koch, 1875) Hottentotta minax occidentalis (Vachon and Stockmann, 1968) Iomachus laeviceps laeviceps (Pocock, 1890) Iomachus politus occidentalis Lourenço, 2003 Iomachus politus politus Pocock, 1896 Kochius bruneus bruneus (Williams, 1970) Lychas variatus canopensis Lourenço and Qi, 2007 Lychas variatus variatus (Thorell, 1876) Microcharmus pauliani ambre Lourenço, Goodman and Fisher, 2006 Microcharmus pauliani namoroka Lourenço, Goodman and Fisher, 2006 Microcharmus pauliani pauliani (Lourenço, 2004) Oiclus purvesii purvesii (Becker, 1880) Oiclus purvesii sabae Francke, 1978 Opisthacanthus africanus africanus Simon, 1876 Opisthacanthus africanus pallidus Lourenço, 2003 Opistophthalmus latimanus latimanus C. L. Koch, 1841 Orthochirus scrobiculosus concolor (Birula, 1898) Orthochirus scrobiculosus dentatus (Birula, 1900) Orthochirus scrobiculosus negebensis (Shulov and Amitai, 1960) Orthochirus scrobiculosus scrobiculosus (Grube, 1873) Pandinus imperator imperator (C. L. Koch, 1841) Pandinus imperator subtypicus (Kraepelin, 1894) Parabuthus granimanus granimanus Pocock, 1895 Parabuthus heterurus heterurus Pocock, 1899 Parabuthus heterurus stefaninii Caporiacco, 1927 Parabuthus mossambicensis flavidus Pocock, 1899 Parabuthus mossambicensis mossambicensis (Peters, 1861) Parabuthus planicauda frenchi (Purcell, 1901) Parabuthus planicauda planicauda (Pocock, 1889) Paruroctonus arenicola arenicola Haradon, 1984 Paruroctonus arenicola nudipes Haradon, 1984 Paruroctonus bantai bantai (Gertsch and Soleglad, 1966) Paruroctonus bantai saratoga Haradon, 1985 Paruroctonus borregoensis actites Haradon, 1984 Paruroctonus borregoensis borregoensis Williams, 1972 Paruroctonus shulovi nevadae Haradon, 1985 Paruroctonus shulovi shulovi (Williams, 1970) Sassanidothus zarudnyi sarghadensis (Birula, 1903) Sassanidothus zarudnyi zarudnyi (Birula, 1900) Scorpio maurus arabicus (Pocock, 1900) Scorpio maurus behringsi Schenkel, 1949 Scorpio maurus legionis Werner, 1932 Scorpio maurus maurus Linnaeus, 1758 Scorpio maurus stemmleri Schenkel, 1949 Scorpio maurus townsendi (Pocock, 1900) Scorpio maurus trarasensis Bouisset and Larrouy, 1962 Scorpio maurus tunetanus Birula, 1910 Scorpio maurus yemenensis Werner, 1936 Scorpiops hardwickii hardwickii (Gervais, 1843) Serradigitus gertschi gertschi (Williams, 1968) Serradigitus gertschi striatus (Hjelle, 1972) Smeringurus vachoni immanis (Soleglad, 1972) Smeringurus vachoni vachoni (Stahnke, 1961) Thestylus glasioui glasioui (Bertkau, 1880) Thestylus glasioui lineatus Giltay, 1928 Tityus argentinus argentinus Borelli, 1899 Tityus argentinus scaber (Werner, 1939) Tityus bahiensis bahiensis (Perty, 1833) Tityus bahiensis eickstedtae Lourenço, 1982 Tityus confluens bodoquena Lourenço, Cabral and Ramos, 2004 Tityus confluens confluens Borelli, 1899 Tityus dasyurus dasyurus Pocock, 1897 Tityus dasyurus fulvipes Mello-Leitão, 1945 Uroctonus mordax mordax Thorell, 1876 Uroctonus mordax pluridens Hjelle, 1972 Urophonius brachycentrus brachycentrus (Thorell, 1876) Uroplectes carinatus australis Hewitt, 1918 Uroplectes carinatus carinatus (Pocock, 1890) Uroplectes formosus basuticus Hewitt, 1927 Uroplectes formosus formosus Pocock, 1890 Uroplectes formosus maculipes Hewitt, 1918 Uroplectes formosus spenceri Pocock, 1896 Uroplectes occidentalis monardi Vachon, 1950 Uroplectes occidentalis occidentalis Simon, 1876 Uroplectes otjimbinguensis massacarum Monard, 1937 Uroplectes otjimbinguensis otjimbinguensis (Karsch, 1879) Uroplectes planimanus planimanus (Karsch, 1879)
Allopalaeophonus Palaeophonus hunteriсилура Шкотске[ 3]
Palaeophonus nuncius силурски фосили из ШведскеФосили шкорпиона пронађени су у многим слојевима , укључујући морске силуријске и естуарске девонске наслаге, наслаге угља из периода карбона и у ћилибару . Да ли су рани шкорпиони били морски или копнени, расправљало се, иако су имали плућа као што су модерне копнене врсте.[ 4] [ 5] [ 6] [ 7] [ 8] Dolichophonus loudonensis [ 9] Gondwanascorpio Гондвана .[ 10] композитне очи сличне онима еуриптерида.[ 11] тријаса Protochactas Protobuthus Chactoidea и Buthoidea , што указује да се крунска група модерних шкорпиона појавила до тог времена.[ 12]
^ а б „Шкорпије” . Бионет Школа. Архивирано из оригинала 08. 03. 2022. г. Приступљено 05. 03. 2021 . ^ „Правилник о проглашењу и заштити строго заштићених и заштићених дивљих врста биљака, животиња и гљива - Прилог II Заштићене врсте које се налазе на Додацима CITES конвенције и на Додатку B Уредбе Комисије (ЕЗ) број 750/2013 од 29. јула 2013. године која допуњује Уредбу Савета (ЕЗ) број 338/97 о заштити врста дивље фауне и флоре регулисањем њихове трговине” . Службени гласник РС. ^ Pocock, R. I. (1901). „The Scottish Silurian Scorpion ”. The Quarterly Journal of Microscopical Science . 44 . Plate 19 — преко Викизворника . ^ Howard, Richard J.; Edgecombe, Gregory D.; Legg, David A.; Pisani, Davide; Lozano-Fernandez, Jesus (2019). „Exploring the Evolution and Terrestrialization of Scorpions (Arachnida: Scorpiones) with Rocks and Clocks”. Organisms Diversity & Evolution . 19 (1): 71—86. ISSN 1439-6092 . doi :10.1007/s13127-019-00390-7 ^ Scholtz, Gerhard; Kamenz, Carsten (2006). „The Book Lungs of Scorpiones and Tetrapulmonata (Chelicerata, Arachnida): Evidence for Homology and a Single Terrestrialisation Event of a Common Arachnid Ancestor”. Zoology . 109 (1): 2—13. PMID 16386884 . doi :10.1016/j.zool.2005.06.003 . ^ Dunlop, Jason A.; Tetlie, O. Erik; Prendini, Lorenzo (2008). „Reinterpretation of the Silurian Scorpion Proscorpius osborni (Whitfield): Integrating Data from Palaeozoic and Recent Scorpions”. Palaeontology . 51 (2): 303—320. doi :10.1111/j.1475-4983.2007.00749.x ^ Kühl, G.; Bergmann, A.; Dunlop, J.; Garwood, R. J.; Rust, J. (2012). „Redescription and Palaeobiology of Palaeoscorpius devonicus Lehmann, 1944 from the Lower Devonian Hunsrück Slate of Germany”. Palaeontology . 55 (4): 775—787. doi :10.1111/j.1475-4983.2012.01152.x ^ Dunlop, J. A.; Penney, D. (2012). Fossil Arachnids . Siri Scientific Press. стр. 23. ISBN 978-0956779540 . ^ Anderson, Evan P.; Schiffbauer, James D.; Jacquet, Sarah M.; Lamsdell, James C.; Kluessendorf, Joanne; Mikulic, Donald G. (2021). „Stranger than a scorpion: a reassessment of Parioscorpio venator, a problematic arthropod from the Llandoverian Waukesha Lagerstätte” . Palaeontology (на језику: енглески). 64 (3): 429—474. ISSN 1475-4983 . doi :10.1111/pala.12534 . Архивирано из оригинала 20. 4. 2021. г. Приступљено 20. 4. 2021 . ^ Gess, R. W. (2013). „The Earliest Record of Terrestrial Animals in Gondwana: a Scorpion from the Famennian (Late Devonian) Witpoort Formation of South Africa” . African Invertebrates 54 (2): 373—379. doi :10.5733/afin.054.0206 Архивирано из оригинала 6. 9. 2013. г. Приступљено 28. 8. 2013 . ^ Insights into the 400 million-year-old eyes of giant sea scorpions (Eurypterida) suggest the structure of Palaeozoic compound eyes ^ Magnani, Fabio; Stockar, Rudolf; Lourenço, Wilson R. (2022). Lionel Delaunay. „Une nouvelle famille, genre et espèce de scorpion fossile du Calcaire de Meride (Trias Moyen) du Mont San Giorgio (Suisse)A new family, genus and species of fossil scorpion from the Meride Limestone (Middle Triassic) of Monte San Giorgio (Switzerland)” . Faunitaxys (на језику: енглески). 10 (24): 1—7. doi :10.57800/FAUNITAXYS-10(24) .
Догељ, В, А: Зоологија бескичмењака, Научна књига , Београд, 1971.
Крунић, М: Зоологија инвертебрата 1, Научна књига , Београд, 1977.
Крунић, М: Зоологија инвертебрата 2, Научна књига , Београд
Маричек, Магдалена, Ћурчић, Б, Радовић, И: Специјална зоологија, Научна књига , Београд, 1986.
Матоничкин, И, Хабдија, И, Примц - Хабдија, Б: Бескраљешњаци - билогија нижих авертебрата, Школска књига , Загреб, 1998.
Marcon, E, Mongini, M: Све животиње света, ИРО Вук Караџић , Београд, 1986.
Петров, И: Сакупљање, препаровање и чување инсеката у збиркама, Биолошки факултет, Београд, 2000.
Радовић, И, Петров, Бригита: Разноврсност живота 1 - структура и функција, Биолошки факултет Београд и Stylos Нови Сад, Београд, 2001.
Polis, Gary (1990). The Biology of Scorpions . Stanford University Press . ISBN 978-0-8047-1249-1 . OCLC 18991506 . Stockmann, Roland; Ythier, Eric (2010). Scorpions of the World . N. A. P. Editions. ISBN 978-2913688117 . Stockmann, Roland (2015). „Introduction to Scorpion Biology and Ecology”. Ур.: Gopalakrishnakone, P.; Possani, L.; F. Schwartz, E.; Rodríguez de la Vega, R. Scorpion Venoms . Springer . стр. 25—59. ISBN 978-94-007-6403-3 . „Scorpion” . The American Heritage Dictionary of the English Language , 5th EditionАрхивирано из оригинала 22. 11. 2020. г. Приступљено 7. 12. 2020 .
